Pollen Specialist Bees of the Eastern United States

Jarrod Fowler & Sam Droege

Pollen Specialist Bee Phenology: OCTOBER

Table 1. Pollen specialist bee families (n = 5), genera (n = 11), and species (n = 48) with taxonomic authorities, distributions according to records from 20 states (AL, CT, DE, FL, GA, LA, MA, MD, ME, MS, NC, NH, NJ, NY, PA, RI, SC, VA, VT, and WV), regional conservation status (Common, Uncommon, or Rare), phenology according to records from 12 months (January, February, March, April, May, June, July, August, September, October, November, and December), and host plant families, tribes, and genera. Bee families and member genera and species are presented in ascending alphabetical order; families, subfamilies, tribes, and subtribes link to BugGuide profiles; species link to Discover Life species profiles. Month headings link to month-specific lists. Host plants link to USDA PLANTS Database genera profiles: view 'Subordinate Taxa' tabs for regional host plant species. Arthropoda: Hexapoda: Insecta: Hymenoptera: Aculeata: Anthophila (Apoidea):

Family: Subfamily: Tribe: Subtribe	Genus (Subgenus) species	Authority	Recorded states	Status	J	F	M	A	M	J	J	A	S	O	N	D	Host plant Family: Tribe: Genera
Andrenidae: Andreninae	Andrena (Callandrena s.l.) accepta	Viereck, 1916	FL-LA-MD-MS-NC-NJ-NY-SC-VA	Rare	.	.	.	.	.	.	.	X	X	X	.	.	Asteraceae: Grindelia Willd., Helianthus L.
	Andrena (Callandrena s.l.) asteroides	Mitchell, 1960	AL-GA-MD-MS-NC-NJ-PA-SC-VA	Rare	.	.	.	.	.	.	.	.	X	X	X	.	Symphyotrichum Nees
	Andrena (Callandrena s.l.) braccata	Viereck, 1907	CT-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA	Common-Rare	.	.	.	.	.	.	.	X	X	X	.	.	Euthamia Nutt. ex Cass., Solidago L.
	Andrena (Callandrena) duplicata	Mitchell, 1960	CT-NC-NJ-NY	Rare	.	.	.	.	.	.	.	X	X	X	.	.	Asteraceae: Bidens L.
	Andrena (Callandrena s.l.) fulvipennis	Smith, 1853	AL-DE-FL-GA-MD-NC-NJ-NY-SC	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
	Andrena (Cnemidandrena) hirticincta	Provancher, 1888	CT-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA-VT-WV	Common	.	.	.	.	.	.	.	X	X	X	.	.	Euthamia Nutt ex. Cass., Solidago L., Symphyotrichum Nees
	Andrena (Cnemidandrena) nubecula	Smith, 1853	CT-DE-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA-VT-WV	Common-Uncommon	.	.	.	.	.	.	.	X	X	X	.	.	Euthamia Nutt. ex Cass., Solidago L., Symphyotrichum Nees
	Andrena (Callandrena s.l.) placata	Mitchell, 1960	CT-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA-VT-WV	Common	.	.	.	.	.	.	X	X	X	X	.	.	Asteraceae: Astereae: Eurybia (Cass.) Cass., Solidago L., Symphyotrichum Nees
	Andrena (Callandrena s.l.) simplex	Smith, 1853	AL-CT-DE-GA-LA-MA-MD-NC-NH-NJ-NY-PA-RI-VA-WV	Common	.	.	.	.	.	.	X	X	X	X	X	.	Solidago L., Symphyotrichum Nees
Andrenidae: Panurginae: Calliopsini	Calliopsis (Calliopsima) coloradensis	Cresson, 1878	AL-GA-LA-MS-NC	Uncommon-Rare	.	.	.	.	.	X	X	X	X	X	.	.	Asteraceae: Chrysopsis (Nutt.) Elliot, Grindelia Willd., Solidago L., Symphyotrichum Nees
Andrenidae: Panurginae: Panurgini: Perditina	Perdita (Cockerellia) albipennis	Cresson, 1868	MS	Rare	.	.	.	.	.	X	X	X	X	X	.	.	Asteraceae: Grindelia Willd., Helianthus L., Heterotheca Cass., Rudbeckia L., Symphyotrichum Nees, Verbesina L.
	Perdita (Hexaperdita) boltoniae	(Robertson, 1902)	DE-FL-GA-LA-MD-MS-NC-NJ-NY-SC-VA	Rare	.	.	.	.	.	.	X	X	X	X	.	.	Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
	Perdita (Perdita) consobrina	Timberlake, 1928	AL-FL-NC-SC-VA	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Asteraceae: Balduina Nutt., Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt., Solidago L.
	Perdita (Perdita) discreta	Timberlake, 1954	NC-SC	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Asteraceae: Astereae: Heterotheca Cass.
	Perdita (Perdita) georgica	Timberlake, 1928	FL-GA-MS-NC	Uncommon	.	.	.	.	.	.	.	X	X	X	.	.	Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
	Perdita (Perdita) gerardiae	Crawford, 1932	FL-MD-MS-NC-NJ-SC	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Agalinis Raf.
	Perdita (Hexaperdita) graenicheri	Timberlake, 1947	FL	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
	Perdita (Perdita) octomaculata	(Say, 1824)	AL-CT-DE-FL-GA-MA-MD-ME-MS-NC-NH-NJ-NY-RI-SC-VA-VT-WV	Common-Uncommon	.	.	.	.	.	.	X	X	X	X	.	.	Asteraceae: Astereae: Baccharis L., Chrysopsis (Nutt.) Elliot, Euthamia Nutt. ex Cass., Grindelia Willd., Pityopsis Nutt., Solidago L., Symphyotrichum Nees
	Perdita (Perdita) polygonellae	Timberlake, 1954	AL-FL-NC	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Polygonella Michx.
Andrenidae: Panurginae: Protandrenini	Pseudopanurgus compositarum	(Robertson, 1893)	CT-DE-GA-LA-MD-NC-NJ-NY-PA-VA-WV	Uncommon	.	.	.	.	.	.	.	.	X	X	.	.	Asteraceae: Bidens L., Rudbeckia L., Solidago L., Symphyotrichum Nees
	Pseudopanurgus labrosiformis	(Robertson, 1898)	GA-MD-NC-VA-WV	Rare	.	.	.	.	.	.	.	X	X	X	.	.	Asteraceae: Bidens L., Coreopsis L., Helianthus L., Rudbeckia L., Silphium L., Solidago L.
	Pseudopanurgus solidaginis	(Robertson, 1893)	AL-MD-MS-NC	Uncommon	.	.	.	.	.	.	.	X	X	X	.	.	Asteraceae: Bidens L., Coreopsis L., Helianthus L., Pityopsis Nutt., Rudbeckia L., Solidago L.
Apidae: Apinae: Eucerini	Melissodes (Eumelissodes) agilis	Cresson, 1878	AL-CT-DE-FL-GA-LA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Common-Uncommon	.	.	.	.	X	X	X	X	X	X	X	.	Asteraceae: Grindelia Willd., Helianthus L., Ratibida Raf., Solidago L.
	Melissodes (Eumelissodes) boltoniae	Robertson, 1905	AL-DE-FL-GA-LA-MD-MS-NC-NY-PA-SC-VA	Uncommon	.	.	.	.	.	.	X	X	X	X	X	.	Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Cirsium Mill., Coreopsis L., Grindelia Willd., Helianthus L., Heterotheca Cass., Ratibida Raf., Rudbeckia L., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
	Melissodes (Eumelissodes) denticulatus	Smith, 1854	AL-CT-DE-FL-GA-LA-MA-MD-MS-NC-NJ-NY-PA-VA-VT-WV	Common-Uncommon	.	.	.	.	X	X	X	X	X	X	.	.	Asteraceae: Vernonia Schreb.
	Melissodes (Eumelissodes) dentiventris	Smith, 1854	AL-CT-DE-FL-GA-MA-MD-MS-NC-NJ-NY-PA-SC-VA	Uncommon	.	.	.	.	.	.	X	X	X	X	.	.	Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Coreopsis L., Helianthus L., Pityopsis Nutt., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
	Melissodes (Heliomelissodes) desponsus	Smith, 1854	AL-CT-DE-FL-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Common	.	.	.	.	.	X	X	X	X	X	.	.	Asteraceae: Cirsium Mill.
	Melissodes (Eumelissodes) druriellus	(Kirby, 1802)	AL-CT-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Uncommon	.	.	.	.	X	X	X	X	X	X	X	.	Asteraceae: Euthamia Nutt. ex Cass., Helianthus L., Rudbeckia L., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
	Melissodes (Eumelissodes) fumosus	LaBerge, 1961	MD-MS-NC-SC-VA	Rare	.	.	.	.	.	X	X	X	X	X	.	.	Asteraceae: Chrysopsis (Nutt.) Elliot, Pityopsis Nutt., Solidago L.
	Melissodes (Eumelissodes) niveus	Robertson, 1895	AL-MD-MS-NC-NJ-NY-VA	Uncommon	.	.	.	.	.	.	X	X	X	X	.	.	Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Helianthus L., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
	Melissodes (Eumelissodes) pilleatus	LaBerge, 1961	NC	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Asteraceae: Chrysopsis (Nutt.) Elliot, Symphyotrichum Nees
	Melissodes (Eumelissodes) tinctus	LaBerge, 1961	FL-GA-LA-MS-NC-SC	Rare	.	.	.	.	.	.	.	X	X	X	X	X	Asteraceae: Chrysopsis (Nutt.) Elliot, Helianthus L., Symphyotrichum Nees, Verbesina L.
	Melissodes (Eumelissodes) trinodis	Robertson, 1901	CT-DE-FL-GA-LA-MA-MD-ME-MS-NC-NJ-NY-RI-VA-VT	Uncommon	.	.	.	.	.	.	X	X	X	X	.	.	Asteraceae: Helianthus L., Ratibida Raf., Rudbeckia L.
Apidae: Apinae: Eucerini	Svastra (Epimelissodes) aegis	(LaBerge, 1956)	AL-FL-GA-LA-MS-NC-SC	Uncommon	.	.	.	.	.	.	X	X	X	X	.	.	Asteraceae: Balduina Nutt., Chrysopsis (Nutt.) Elliot, Helenium L., Helianthus L., Symphyotrichum Nees
	Svastra (Anthedonia) compta	(Cresson, 1878)	GA-MD-NC-NJ-PA	Uncommon	.	.	.	.	.	X	X	X	X	X	X	.	Oenothera L.
	Svastra (Epimelissodes) obliqua	(Say, 1837)	AL-DE-FL-GA-LA-MD-MS-NC-NJ-NY-SC-VA	Uncommon	.	.	.	.	.	X	X	X	X	X	X	.	Asteraceae: Cirsium Mill., Grindelia Willd., Helianthus L., Rudbeckia L., Vernonia Schreb.
Apidae: Apinae: Exomalopsini	Anthophorula (Anthophorisca) micheneri	Timberlake, 1948	MS-VA	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Agalinis Raf.
Colletidae	Colletes americanus	Cresson, 1868	CT-FL-GA-LA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-VA-WV	Common-Uncommon	.	.	.	.	X	X	X	X	X	X	.	.	Asteraceae: Euthamia Nutt. ex Cass., Grindelia Willd., Helianthus L., Pityopsis Nutt., Rudbeckia L., Solidago L., Symphyotrichum Nees
	Colletes ciliatus	Patton, 1879	MD-VA	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Cuscuta L.
	Colletes compactus	Cresson, 1868	CT-DE-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA-VT-WV	Common	.	.	.	.	.	X	X	X	X	X	X	.	Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Rudbeckia L., Solidago L., Symphyotrichum Nees
	Colletes howardi	Swenk, 1925	FL-NC	Rare	.	.	.	.	.	.	.	.	X	X	.	.	Dalea L.
	Colletes simulans	Cresson, 1868	AL-CT-FL-GA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Common	.	.	.	.	X	X	X	X	X	X	X	.	Euthamia Nutt. ex Cass., Solidago L., Symphyotrichum Nees
	Colletes speculiferus	Cockerell, 1927	CT-DE-FL-GA-MA-MD-MS-NC-NJ-NY-RI-SC-VA	Uncommon	.	.	.	.	.	.	X	X	X	X	.	.	Asteraceae: Euthamia Nutt ex. Cass., Solidago L., Symphyotrichum Nees
	Colletes thysanellae	Mitchell, 1951	AL-FL-GA-NC-SC-VA	Rare	.	.	.	.	.	.	.	.	X	X	X	.	Asteraceae: Baccharis L., Balduina Nutt., Heterotheca Cass., Solidago L.
Megachilidae: Anthidiini	Trachusa (Heteranthidium) zebrata	(Cresson, 1872)	LA-MS	Rare	.	.	.	.	.	.	.	X	X	X	.	.	Asteraceae: Helianthus L.
Megachilidae: Megachilini	Megachile (Sayapis) inimica	Cresson, 1872	AL-CT-DE-FL-GA-LA-MA-MD-MS-NC-NJ-NY-PA-SC-VA-WV	Uncommon	.	.	.	.	.	.	X	X	X	X	X	.	Asteraceae: Baccharis L., Chrysopsis (Nutt.) Elliot, Cirsium Mill., Coreopsis L., Erigeron L., Grindelia Willd., Helianthus L., Rudbeckia L., Silphium L., Solidago L., Verbesina L., Vernonia Schreb.
	Megachile (Melanosarus) xylocopoides	Smith, 1853	AL-DE-FL-GA-LA-MD-MS-NC-NJ-SC-VA	Uncommon	.	.	X	X	X	X	X	X	X	X	.	.	Asteraceae: Coreopsis L., Erigeron L., Helianthus L., Rudbeckia L., Solidago L., Vernonia Schreb.
Melittidae: Dasypodainae: Hesperapini	Hesperapis (Carinapis) oraria	Snelling & Stage, 1997	AL-FL-MS	Rare	.	.	.	.	.	.	.	.	.	X	.	.	Asteraceae: Balduina Nutt.

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