Pollen Specialist Bees of the Eastern United States

Jarrod Fowler & Sam Droege

Pollen Specialist Bee Phenology: MAY

Table 1. Pollen specialist bee families (n = 6), genera (n = 22), and species (n = 89) with taxonomic authorities, distributions according to records from 20 states (AL, CT, DE, FL, GA, LA, MA, MD, ME, MS, NC, NH, NJ, NY, PA, RI, SC, VA, VT, and WV), regional conservation status (Common, Uncommon, or Rare), phenology according to records from 12 months (January, February, March, April, May, June, July, August, September, October, November, and December), and host plant families, tribes, and genera. Bee families and member genera and species are presented in ascending alphabetical order; families, subfamilies, tribes, and subtribes link to BugGuide profiles; species link to Discover Life species profiles. Month headings link to month-specific lists. Host plants link to USDA PLANTS Database genera profiles: view 'Subordinate Taxa' tabs for regional host plant species. Arthropoda: Hexapoda: Insecta: Hymenoptera: Aculeata: Anthophila (Apoidea):

Family: Subfamily: Tribe: Subtribe	Genus (Subgenus) species	Authority	Recorded states	Status	J	F	M	A	M	J	J	A	S	O	N	D	Host plant Family: Tribe: Genera
Andrenidae: Andreninae	Andrena (Parandrena) andrenoides	(Cresson, 1878)	AL-FL-GA-MD-NC-NJ-NY-SC-VA	Rare	.	.	.	X	X	.	.	.	.	.	.	.	Salix L.
	Andrena (Scaphandrena) arabis	Robertson, 1897	CT-DE-MA-MD-NC-NJ-NY-PA-VA-WV	Uncommon	.	.	X	X	X	.	.	.	.	.	.	.	Arabis L., Cardamine L.
	Andrena (Thysandrena) bisalicis	Viereck, 1908	AL-CT-DE-GA-LA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Common-Uncommon	.	.	X	X	X	X	X	.	.	.	.	.	Salix L.
	Andrena (Conandrena) bradleyi	Viereck, 1907	CT-DE-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-SC-VA	Common-Uncommon	.	.	X	X	X	X	.	.	.	.	.	.	Ericaceae: Chamaedaphne Moench, Kalmia L., Vaccinium L.
	Andrena (Andrena) carolina	Viereck, 1909	AL-GA-CT-DE-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA-WV	Common-Uncommon	.	.	.	X	X	X	X	.	.	.	.	.	Ericaceae: Gaylussacia Kunth, Vaccinium L.
	Andrena (Andrena) clarkella	(Kirby, 1802)	CT-MA-ME-NH-NY-RI-VT-WV	Common-Rare	.	.	.	X	X	X	X	.	.	.	.	.	Salix L.
	Andrena (Andrena) cornelli	Viereck, 1907	CT-DE-GA-MA-MD-NC-NJ-NY-PA-VA	Uncommon	.	.	.	.	X	X	X	.	.	.	.	.	Rhododendron L.
	Andrena (Ptilandrena) distans	Provancher, 1888	CT-GA-MA-MD-NC-NH-NJ-NY-PA-RI-VA	Common-Uncommon	.	.	.	X	X	X	.	.	.	.	.	.	Geranium L.
	Andrena (Ptilandrena) erigeniae	Robertson, 1891	CT-DE-MA-MD-MS-NC-NH-NJ-NY-PA-SC-VA-WV	Common	.	.	X	X	X	.	.	.	.	.	.	.	Claytonia L.
	Andrena (Tylandrena) erythrogaster	(Ashmead, 1890)	CT-FL-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Common-Uncommon	.	.	.	X	X	X	X	.	.	.	.	.	Salix L.
	Andrena (Leucandrena) erythronii	Robertson, 1891	CT-FL-MA-MD-ME-NH-NJ-NY-PA-VA-WV	Uncommon	.	.	.	X	X	X	.	.	.	.	.	.	Erythronium L.
	Andrena (Gonandrena) fragilis	Smith, 1853	CT-DE-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-SC-VA-WV	Common	.	.	.	.	X	X	X	.	.	.	.	.	Cornus (Swida) L.
	Andrena (Andrena) frigida	Smith, 1853	CT-GA-MA-MD-ME-NH-NJ-NY-PA-RI-VA	Common-Uncommon	.	.	X	X	X	X	.	.	.	.	.	.	Salix L.
	Andrena (Callandrena s.l.) gardineri	Cockerell, 1906	AL-FL-GA-MD-MS-NC-SC-VA-WV	Uncommon	.	.	.	X	X	X	.	.	.	.	.	.	Packera Á. Löve & D. Löve
	Andrena (Euandrena) geranii	Robertson, 1891	FL-MD-NH-NJ-NY-PA-VA-VT-WV	Uncommon	.	.	.	X	X	X	X	.	.	.	.	.	Hydrophyllum L.
	Andrena (Micrandrena) illinoiensis	Robertson, 1891	MD-NY-VA	Rare	.	.	.	X	X	X	.	.	.	.	.	.	Salix L.
	Andrena (Gonandrena) integra	Smith, 1853	CT-MA-ME-MD-NC-NJ-NY-PA-VA-VT	Uncommon	.	.	.	.	X	X	X	X	X	.	.	.	Cornus (Swida) L.
	Andrena (Scrapteropsis) kalmiae	Atwood, 1934	CT-MA-ME-NH	Uncommon	.	.	.	X	X	X	X	.	.	.	.	.	Ericaceae: Kalmia L., Lyonia Nutt., Vaccinium L.
	Andrena (Callandrena s.l.) krigiana	Robertson, 1901	AL-CT-FL-GA-LA-MA-MD-MS-NC-NH-NJ-NY-PA-SC-VA-WV	Uncommon	.	.	X	X	X	X	.	.	.	.	.	.	Asteraceae: Cichorieae: Hieracium L., Krigia Schreb.
	Andrena (Micrandrena) lamelliterga	Ribble, 1968	MD-PA-VA	Rare	.	.	.	X	X	.	.	.	.	.	.	.	Phacelia Juss.
	Andrena (Andrena) macoupinensis	Robertson, 1900	AL-FL-GA-MD-MS-NC-PA-SC-VA-WV	Uncommon	.	.	X	X	X	X	.	.	.	.	.	.	Salix L.
	Andrena (Trachandrena) mariae	Robertson, 1891	GA-MD-NC-NH-NJ-RI-VA-WV	Uncommon	.	.	X	X	X	.	.	.	.	.	.	.	Salix L.
	Andrena (Micrandrena) melanochroa	Cockerell, 1898	CT-GA-MA-MD-ME-NC-NH-NJ-NY-VA-WV	Uncommon	.	.	X	X	X	X	X	.	.	.	.	.	Rosaceae: Fragaria L., Potentilla L., Rubus L.
	Andrena (Parandrena) nida	Mitchell, 1960	AL-FL-GA-MD-MS-NC-NH-NJ-NY-RI-VA-WV	Uncommon	.	.	X	X	X	.	.	.	.	.	.	.	Salix L.
	Andrena (Micrandrena) nigrae	Robertson, 1905	AL-CT-DE-FL-GA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Uncommon	.	.	.	X	X	X	.	.	.	.	.	.	Salix L.
	Andrena (Gonandrena) perisimulata	Viereck, 1917	CT-MA-ME-NH-NY	Uncommon	.	.	.	.	X	X	X	.	.	.	.	.	Cornus (Swida) L.
	Andrena (Euandrena) phaceliae	Mitchell, 1960	MD-NC-VA-WV	Rare	.	.	.	X	X	.	.	.	.	.	.	.	Phacelia Juss.
	Andrena (Euandrena) polemonii	Robertson, 1891	DE-WV	Rare	.	.	.	.	X	.	.	.	.	.	.	.	Polemonium L.
	Andrena (Trachandrena) rehni	Viereck, 1907	CT, DE, GA, MA, MD, NC, NJ, NY, PA, VA	Rare	.	.	.	X	X	X	.	.	.	.	.	.	Castanea Mill.
	Andrena (Callandrena s.l.) rudbeckiae	Robertson, 1891	DE-FL-LA-MD-MS-NC-NJ-NY-PA	Rare	.	.	.	.	X	X	X	X	.	.	.	.	Ratibida Raf., Rudbeckia L.
	Andrena (Micrandrena) salictaria	Robertson, 1905	CT-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA-VT-WV	Common-Rare	.	.	X	X	X	.	.	.	.	.	.	.	Salix L.
	Andrena (Trachandrena) sigmundi	Cockerell, 1902	CT-GA-MA-ME-NC-NH-RI-VT	Common	.	.	X	X	X	X	.	.	.	.	.	.	Salix L.
	Andrena (Derandrena) uvulariae	Mitchell, 1960	DE-MA-MD-NC-NH-NJ-NY-PA-SC-VA-WV	Rare	.	.	.	.	X	.	.	.	.	.	.	.	Uvularia L.
	Andrena (Parandrena) wellesleyana	Robertson, 1897	CT-MA-RI	Uncommon	.	.	.	X	X	.	.	.	.	.	.	.	Salix L.
	Andrena (Micrandrena) ziziae	Robertson, 1891	CT-GA-MA-MD-NC-NH-NJ-NY-PA-VA-VT-WV	Common-Uncommon	.	.	.	X	X	X	.	.	.	.	.	.	Zizia W.D.J. Koch
	Andrena (Derandrena) ziziaeformis	Cockerell, 1908	CT-DE-GA-MD-NC-NJ-NY-PA-VA-WV	Uncommon-Rare	.	.	.	X	X	.	.	.	.	.	.	.	Potentilla L., Waldsteinia Willd.
Andrenidae: Panurginae: Panurgini: Panurginia	Panurginus atramontensis	Crawford, 1926	GA-MD-MS-NC-NJ-VA	Rare	.	.	.	X	X	X	.	.	.	.	.	.	Vaccinium (Polycodium) L.
	Panurginus potentillae	(Crawford, 1916)	CT-GA-MA-MD-NC-NJ-NY-SC-VA	Uncommon-Rare	.	.	X	X	X	.	.	.	.	.	.	.	Potentilla L.
Andrenidae: Panurginae: Panurgini: Perditina	Perdita (Alloperdita) floridensis	Timberlake, 1928	FL-NC	Rare	.	.	.	X	X	.	.	.	.	.	.	.	Ilex L.
	Perdita (Hexaperdita) foveata	Timberlake, 1956	FL	Rare	.	.	.	.	X	X	.	.	.	.	.	.	Asteraceae: Coreopsis L.
	Perdita (Perdita) gerhardi	Viereck, 1904	FL-MD-NC-NJ-VA-WV	Rare	.	.	.	X	X	X	X	X	.	.	.	.	Monarda L.
	Perdita (Perdita) halictoides	Smith, 1853	FL-GA-MA-MD-NC-NH-NJ-NY-VT	Uncommon-Rare	.	.	.	X	X	X	X	X	X	.	.	.	Physalis L.
	Perdita (Perdita) maculigera	Cockerell, 1896	NY	Rare	.	.	.	X	X	X	.	.	.	.	.	.	Salix L.
	Perdita (Alloperdita) novaeangliae	Viereck, 1907	FL-CT-MA-NC-RI-SC	Uncommon	.	.	.	.	X	X	X	.	.	.	.	.	Lyonia Nutt.
	Perdita (Hexaperdita) nubila	Timberlake, 1958	FL	Rare	.	.	.	.	X	.	.	.	.	.	.	.	Asteraceae: Erigeron L.
	Perdita (Alloperdita) obscurata	Cresson, 1878	AL-FL-GA-MS-NC	Uncommon-Rare	.	.	.	.	X	.	.	.	.	.	.	.	Diospyros L., Vaccinium L.
Andrenidae: Panurginae: Protandrenini	Protandrena abdominalis	(Cresson, 1878)	GA-MD-NC-NJ-PA-VA-WV	Rare	.	.	.	.	X	X	X	X	X	.	.	.	Monarda L.
Andrenidae: Panurginae: Protandrenini	Pseudopanurgus albitarsis	(Cresson, 1872)	GA-NC-VA	Rare	.	.	.	.	X	X	X	X	X	.	.	.	Asteraceae: Coreopsis L., Helianthus L., Ratibida Raf., Rudbeckia L.
	Pseudopanurgus illinoiensis	(Cresson, 1878)	FL-LA-MD-NC-VA	Rare	.	.	.	.	X	.	.	.	.	.	.	.	Asteraceae: Erigeron L., Heterotheca Cass.
	Pseudopanurgus passiflorae	Robertson, 1897	MS-NC-VA	Rare	.	.	.	X	X	X	X	X	X	.	.	.	Passiflora L.
	Pseudopanurgus virginicus	(Cockerell, 1907)	AL-GA-MD-NC-VA-WV	Rare	.	.	.	.	X	X	.	.	.	.	.	.	Ceanothus L., Houstonia L.
Apidae: Apinae: Anthophorini	Habropoda laboriosa	(Fabricius, 1804)	AL-CT-DE-FL-GA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-SC-VA	Common-Uncommon	X	X	X	X	X	.	.	.	.	.	X	X	Cercis L., Vaccinium L.
Apidae: Apinae: Emphorini	Diadasia (Coquillettapis) afflica	(Cresson, 1878)	LA-MS	Rare	.	.	X	X	X	X	.	.	.	.	.	.	Malvaceae: Callirhoe Nutt.
Apidae: Apinae: Eucerini	Florilegus (Florilegus) condignus	(Cresson, 1878)	AL-DE-FL-GA-LA-MD-MS-NC-NJ-SC-VA	Uncommon-Rare	.	.	.	X	X	X	X	X	.	.	.	.	Pontederia L.
Apidae: Apinae: Eucerini	Melissodes (Eumelissodes) agilis	Cresson, 1878	AL-CT-DE-FL-GA-LA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Common-Uncommon	.	.	.	.	X	X	X	X	X	X	X	.	Asteraceae: Grindelia Willd., Helianthus L., Ratibida Raf., Solidago L.
	Melissodes (Apomelissodes) apicatus	Lovell and Cockerell, 1906	CT-DE-FL-GA-MA-MD-ME-NH-NJ-NY	Uncommon-Rare	.	.	X	X	X	X	X	.	.	.	.	.	Pontederia L.
	Melissodes (Eumelissodes) denticulatus	Smith, 1854	AL-CT-DE-FL-GA-LA-MA-MD-MS-NC-NJ-NY-PA-VA-VT-WV	Common-Uncommon	.	.	.	.	X	X	X	X	X	X	.	.	Asteraceae: Vernonia Schreb.
	Melissodes (Eumelissodes) druriellus	(Kirby, 1802)	AL-CT-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Uncommon	.	.	.	.	X	X	X	X	X	X	X	.	Asteraceae: Euthamia Nutt. ex Cass., Helianthus L., Rudbeckia L., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
	Melissodes (Apomelissodes) fimbriatus	Cresson, 1878	MS-NC-VA	Rare	.	.	.	X	X	X	X	.	.	.	.	.	Oenothera L.
	Melissodes (Apomelissodes) mitchelli	LaBerge, 1956	FL-GA-NC-SC	Rare	.	.	.	X	X	.	.	.	.	.	.	.	Opuntia Mill.
Apidae: Apinae: Eucerini	Svastra (Epimelissodes) petulca	(Cresson, 1878)	AL-FL-GA-LA-MS-NC-NJ-SC	Uncommon-Rare	.	.	.	X	X	X	X	X	X	.	.	.	Asteraceae: Coreopsis L., Helenium L., Helianthus L., Pyrrhopappus DC., Ratibida Raf., Rudbeckia L., Verbesina L.
Apidae: Apinae: Eucerini	Xenoglossa (Eoxenoglossa) kansensis	Cockerell, 1905	FL-NC-SC	Rare	.	.	.	.	X	X	X	X	.	.	.	.	Cucurbita L., Ipomoea L.
	Xenoglossa (Eoxenoglossa) strenua	(Cresson, 1878)	AL-FL-GA-LA-MD-MS-NC-SC-VA	Uncommon	.	.	.	.	X	X	X	X	.	.	.	.	Cucurbita L.
Colletidae	Colletes aestivalis	Patton, 1879	DE-GA-MA-MD-MS-NC-NY-VA	Rare	.	.	.	.	X	X	X	.	.	.	.	.	Heuchera L.
	Colletes americanus	Cresson, 1868	CT-FL-GA-LA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-VA-WV	Common-Uncommon	.	.	.	.	X	X	X	X	X	X	.	.	Asteraceae: Euthamia Nutt. ex Cass., Grindelia Willd., Helianthus L., Pityopsis Nutt., Rudbeckia L., Solidago L., Symphyotrichum Nees
	Colletes banksi	Swenk, 1908	CT-FL-GA-LA-NC-NJ-NY-SC-VA	Rare	.	.	.	X	X	X	X	.	.	.	.	.	Ilex L.
	Colletes brevicornis	Robertson, 1897	FL-GA-LA-MD-NC-NJ-SC-VA-WV	Uncommon	.	.	X	X	X	X	.	.	.	.	.	.	Campanula L., Triodanis Raf. ex Greene
	Colletes brimleyi	Mitchell, 1951	FL-GA-NC-NJ-SC	Rare	.	.	X	X	X	X	.	.	.	.	.	.	Ilex L.
	Colletes latitarsis	Robertson, 1891	AL-CT-FL-GA-MA-MD-ME-MS-NC-NJ-NY-PA-SC-VA-WV	Uncommon	.	.	X	X	X	X	X	X	X	.	.	.	Physalis L.
	Colletes productus	Robertson, 1891	AL-CT-FL-GA-LA-MA-MD-NC-NH-NJ-SC-VA-WV	Uncommon-Rare	.	.	.	X	X	X	X	.	.	.	.	.	Lyonia Nutt., Vaccinium L.
	Colletes simulans	Cresson, 1868	AL-CT-FL-GA-MA-MD-ME-MS-NC-NH-NJ-NY-PA-RI-SC-VA-VT-WV	Common	.	.	.	.	X	X	X	X	X	X	X	.	Euthamia Nutt. ex Cass., Solidago L., Symphyotrichum Nees
	Colletes validus	Cresson, 1868	CT-DE-GA-MA-ME-MD-NC-NH-NJ-NY-PA-RI-SC-VA	Uncommon	.	.	X	X	X	X	X	.	.	.	.	.	Ericaceae: Vaccinium L.
	Colletes willistoni	Robertson, 1891	FL-GA-MD-NC-NJ-NY-PA-VA	Rare	.	.	.	X	X	X	X	X	X	.	.	.	Physalis L.
Halictidae: Halictinae: Halictini	Lasioglossum (Hemihalictus) lustrans	(Cockerell, 1897)	AL-DE-FL-GA-LA-MD-MS-NC-SC-VA	Rare	.	.	.	X	X	X	X	X	X	.	.	.	Pyrrhopappus DC.
Halictidae: Rophitinae	Dufourea novaeangliae	(Robertson, 1897)	CT-MA-MD-ME-NH-NJ-NY-PA-RI-VA-VT	Common-Rare	.	.	.	.	X	X	X	.	.	.	.	.	Pontederia L.
Megachilidae: Anthidiini	Dianthidium (Dianthidium) curvatum	(Smith, 1854)	AL-FL-GA-MS-NC-SC	Uncommon	.	.	.	X	X	X	X	X	X	.	.	.	Asteraceae: Coreopsis L., Grindelia Willd., Helenium L., Helianthus L.
Megachilidae: Lithurginae	Lithurgopsis gibbosa	(Smith, 1853)	AL-FL-GA-NC-SC	Uncommon-Rare	.	.	.	X	X	.	.	.	.	.	.	.	Opuntia Mill.
Megachilidae: Megachilini	Megachile (Megachiloides) integra	Cresson, 1878	AL-FL-GA-MD-NC-NJ-SC	Rare	.	.	X	X	X	X	X	X	X	.	.	.	Galactia P. Br., Strophostyles Elliot
	Megachile (Megachiloides) oenotherae	(Mitchell, 1924)	GA-LA-NC-NJ	Rare	.	.	.	X	X	X	.	.	.	.	.	.	Oenothera L.
	Megachile (Melanosarus) xylocopoides	Smith, 1853	AL-DE-FL-GA-LA-MD-MS-NC-NJ-SC-VA	Uncommon	.	.	X	X	X	X	X	X	X	X	.	.	Asteraceae: Coreopsis L., Erigeron L., Helianthus L., Rudbeckia L., Solidago L., Vernonia Schreb.
Megachilidae: Osmiini	Ashmeadiella (Ashmeadiella) bucconis	(Say, 1837)	GA-LA-NC	Rare	.	.	.	.	X	X	X	X	.	.	.	.	Asteraceae: Coreopsis L., Erigeron L., Grindelia Willd., Heterotheca Cass., Helianthus L., Rudbeckia L., Solidago L., Verbesina L.
Megachilidae: Osmiini	Hoplitis (Robertsonella) micheneri	Mitchell, 1962	FL-GA-NC-SC	Uncommon-Rare	.	.	.	X	X	.	.	.	.	.	.	.	Amorpha L.
	Hoplitis (Robertsonella) simplex	(Cresson, 1864)	DE-MD-NC-NJ-SC-VA-WV	Rare	.	.	X	X	X	.	.	.	.	.	.	.	Nemophila Nutt., Phacelia Juss.
Megachilidae: Osmiini	Osmia (Helicosmia) chalybea	Smith, 1853	AL-DE-FL-GA-LA-MD-MS-NC-NJ-NY-VA-WV	Rare	.	.	X	X	X	.	.	.	.	.	.	.	Cirsium Mill.
	Osmia (Melanosmia) distincta	Cresson, 1864	CT-DE-FL-MA-MD-ME-NC-NH-NJ-NY-PA-VA-VT-WV	Uncommon	.	.	X	X	X	X	X	.	.	.	.	.	Penstemon Schmidel
	Osmia (Helicosmia) texana	Cresson, 1872	MD-VA-WV	Rare	.	.	X	X	X	X	X	X	.	.	.	.	Cirsium Mill.
	Osmia (Melanosmia) virga	Sandhouse, 1939	CT-DE-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA-WV	Uncommon	.	.	.	X	X	X	.	.	.	.	.	.	Vaccinium L.
Melittidae: Melittinae: Macropidini	Macropis (Macropis) ciliata	Patton, 1880	CT-GA-MA-MD-ME-NC-NH-NJ-NY-PA-RI-VA	Uncommon-Rare	.	.	.	.	X	X	X	X	.	.	.	.	Lysimachia L.
Melittidae: Melittinae: Melittini	Melitta (Cilissa) eickworti	Snelling & Stage, 1995	GA-MD-NC-NJ-NY-WV	Rare	.	.	.	.	X	X	.	.	.	.	.	.	Vaccinium (Polycodium) L.

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