Pollen Specialist Bees of the Central United States

Jarrod Fowler (2020)

Introduction

This website compiles associations among native pollen specialist bees and native host plants from the Central United States. First, pollen specialist bees are defined and methods are described. Next, a table composed of pollen specialist bees and associated host plants is presented. Last, advice about conserving native pollen specialist bees is provided. Please note that this compilation is incomplete and in progress. Researchers are invited to add information to this website. Email records to: j@jarrodfowler.com

Visit:
Fowler, J. (2020). Pollen Specialist Bees of the Western United States
Fowler, J. & Droege, S. (2020). Pollen Specialist Bees of the Eastern United States

Pollen Specialist Bees

Roughly 30% of the ~1,800 species of bees native to the Central United States are pollen specialists. Pollen specialist bees evolved a continuum of facultative or obligate associations with flowering host plants (Cane & Sipes 2006; Danforth et al. 2019; Hurd et al. 1980; Larkin, Neff, & Simpson 2008; Robertson 1925; Wright 2018). Oligolectic bees or oligoleges specifically associate with one host plant family [a.k.a. mesolectic bees or mesoleges] or a few related genera or species and monolectic bees or monoleges specifically associate with a single host plant genus or species, while polylectic bees or polyleges generically forage unrelated plants (Cane & Sipes 2006; Danforth et al. 2019; Robertson 1925).

Pollen specialist associations can mutually benefit both bees and flowers from improved foraging effectiveness and efficiency, pollen digestibility, and pollination rates. However, foraging restrictions may create greater susceptibility to harm from pollination or pollinization shortages due to habitat degradation, fragmentation, and loss or phenological mismatch (Minckley et al. 1994; Packer et al. 2005; Rafferty et al. 2015). Therefore, contemporary anthropogenic threats in the Central United States potentially imperil native pollen specialist bee species and their endemic, indigenous host plants with population declines and extinctions through loss of species diversity.

Methods

Records of native pollen specialist bees captured or observed on flowers of host plants were compiled from Discover Life (Ascher & Pickering 2020), peer reviewed articles (Bouseman & LaBerge 1978; Brooks & Griswold 1988; Cockerell 1916, 1919; Cresson 1878; Daly 1973; Danforth 1994; Donovan 1977; Griswold 1993; Griswold & Miller 2010; Hurd et al. 1980; LaBerge 1963, 1967, 1969, 1971a, 1971b, 1973, 1977, 1980, 1985, 1986, 1989; LaBerge & Bouseman 1970; LaBerge & Ribble 1972, 1975; Lanham, 1981; Linsley & MacSwain 1958; McGinley 2003; Michener 1939; Minckley et al. 1994; Moldenke 1976, 1979; Neff 2003; Neff & Larkin 2003; Parys et al. 2018; Portman, Neff, & Griswold 2016; Provancher, 1895; Ribble 1974; Robertson 1926, 1928, 1929; Rozen 1958; Snelling 1983; Thorp 1969; Timberlake 1954, 1956, 1958, 1960, 1962, 1964, 1968, 1975, 1980l Wright 2018), technical bulletins (Danforth 1996; Grigarick & Stange 1968; Hurd & Michener 1955; Krombein et al. 1979; LaBerge 1967; Mitchell 1960, 1962; Ribble 1968; Stephen 1954; Thorp & LaBerge 2005; Timberlake 1953), and personal communications.

If bees and plants were indigenous to the Central United States without human intervention, then bees and plants were defined "native". Definitions of bee conservation status were averaged across ranges. If there were on average: between one and 100 records, then bees were defined "Rare"; between one and 500 records, then bees were defined "Uncommon-Rare"; around 500 records, then bees were defined "Uncommon"; between 500 and 1000 records, then bees were defined "Common-Uncommon"; around 1000 or more records, then bees were defined "Common". Conservation status may vary according to species-specific distributions and dispersions. Bee-flower records were compared with state-level plant distributions from the USDA Plants Database (USDA NRCS 2020) and county-level distributions from The Biota of North America Program (Kartesz, BONAP 2015). This study includes only host plant genera that are native and documented as present in at least one county of at least one of the seventeen regional states: Arkansas, Iowa, Illinois, Indiana, Kansas, Kentucky, Michigan, Minnesota, Missouri, Nebraska, North Dakota, Ohio, Oklahoma, South Dakota, Tennessee, Texas, and Wisconsin. Because plant taxonomy has undergone major recent changes, many names have been updated compared to those published in the original literature of bee-flower associations.

Results

Pollen specialist bee species of the Central United States were alphabetically tabulated with associated host plant families in Table 1. Six families, 10 subfamilies, 14 tribes, two subtribes, 45 genera, 102 subgenera, and 471 species of pollen specialist bees were listed. The bee family with the most pollen specialists was Andrenidae (251 spp.), while the most speciose subfamily was Panurginae (140 spp.), tribe was Panurgini (97 spp.), subtribe was Perditina (94 spp.), genus was Andrena (111 spp.), and subgenus was Perdita (51 spp.). The most recurrent authorities (n = 46 authors) were Theodore Dru Allison Cockerell (1866–1948, 274 spp.), Ezra Townsend Cresson (1838–1926, 147 spp.), Philip Hunter Timberlake (1883–1981, 84 spp.), and Charles Robertson (1858–1935, 78 spp.). Most species were first described during years 1878 (73 spp.), 1896 (60 spp.), and 1897 (48 spp.).

The states most inhabited by pollen specialist bees were: Texas (309 spp.; 41% spp.), Kansas (166 spp.; 45% spp.), Nebraska (164 spp.; 39% spp.), Illinois (145 spp.; 34% spp.), North Dakota (128 spp.; 37% spp.), South Dakota (119 spp.; 37% spp.), Indiana (106 spp.; 27% spp.), Missouri (102 spp.; 38% spp.), Michigan (101 spp.; 23% spp.), Wisconsin (98 spp.; 25% spp.), Minnesota (90 spp.; 27% spp.), Oklahoma (89 spp.; 43% spp.), Ohio (83 spp.; 25% spp.), Iowa (78 spp.; 35% spp.), Tennessee (74 spp.; 28% spp.), Arkasas (62 spp.; 34% spp.), and Kentucky (33 spp.; 22% spp.). Of the 471 species of pollen specialist bees, 148 spp. were recorded only in Texas, while three spp. were recorded in all 17 states: Andrena (Micrandrena) nigrae Robertson, 1905, Melissodes (Eumelissodes) agilis Cresson, 1878, and Svastra (Epimelissodes) obliqua (Say, 1837). Overall, 303 bee species were considered "Rare", while 95 spp. were “Uncommon-Rare", 48 spp. were "Uncommon", 17 spp. were “Common-Uncommon", and eight spp. were "Common". The months of activity for pollen specialist bees relative to associated host plant flowering phenology were August (277 spp.), July (247 spp.), June (228 spp.), September (219 spp.), May (204 spp.), April (172 spp.), March (90 spp.), October (88 spp.), November (23 spp.), February (6 spp.), December (6 spp.), and January (1 spp.). Oftentimes, pollen specialist bees had only three month (139 spp.) or two month (113 spp.) flight periods.

The most recurrent host plant families among pollen specialist bee species were Asteraceae (198 bee spp.) and Fabaceae (29 spp.). The top 25 most recurrent host plant genera (N = ~250) associated with pollen specialist bee species were: Helianthus L. (98 spp.), Grindelia Willd. (67 spp.), Heterotheca Cass. (63 spp.), Solidago L. (57 spp.), Chrysothamnus Nutt. (51 spp.), Symphyotrichum Nees (50 spp.), Gutierrezia Lag. (49 spp.), Verbesina L. (48 spp.), Ericameria Nutt. (46 spp.), Baileya Harv. & A. Gray ex A. Gray (36 spp.), Rudbeckia L. (33 spp.), Erigeron L. (32 spp.), Cirsium Mill. (31 spp.), Coreopsis L. (28 spp.), Machaeranthera Nees (23 spp.), Senecio L. (22 spp.), Salix L. (19 spp.), Prosopis L. (17 spp.), Bidens L. (18 spp.), Dalea L. (17 spp.), Isocoma Nutt. (17 spp.), Baccharis L. (14 spp.), Encelia Adans. (16 spp.), and Heliomeris Nutt. (16 spp.).

Tables

Table 1. Pollen specialist bee families (n = 6), genera (n = 45), and species (n = 471) with taxonomic authorities, distributions according to records from 11 states (AR, IA, IL, IN, KS, KY, MI, MN, MO, ND, NE, OH, OK, SD, TN, TX, and WI), regional conservation status (Common, Uncommon, or Rare), phenology according to records from 12 months, and host plant families, tribes, and genera. Bee families and member genera and species are presented in ascending alphabetical order; families, subfamilies, tribes, and subtribes link to BugGuide profiles; species link to Discover Life species profiles. Host plants link to USDA PLANTS Database genera profiles: view 'Subordinate Taxa' tabs for regional host plant species. Arthropoda: Hexapoda: Insecta: Hymenoptera: Aculeata: Anthophila (Apoidea):
Family: Subfamily: Tribe: SubtribeGenus (Subgenus) speciesAuthorityStatesStatusJFMAMJJASONDHost plant Family: Tribe: Genera
Andrenidae: Andreninae Andrena (Callandrena) accepta Viereck, 1916 IA-IL-IN-KS-MO-ND-NE-OK-SD-TN-TX Uncommon ......XXXX.. Asteraceae: Helianthus L.
Andrena (Callandrena s.l.) afimbriata LaBerge, 1967 TX Rare ...X........ Asteraceae: Berlandiera DC., Pyrrhopappus DC.
Andrena (Scrapteropsis) alamonis Viereck, 1917 TX Rare ..XX........ Lesquerella S. Watson
Andrena (Callandrena s.l.) aliciae Robertson, 1891 IA-IL-IN-KS-KY-MI-MN-NE-OH-TN-WI Rare ......XXX... Asteraceae: Bidens L., Helianthus L., Rudbeckia L., Silphium L., Solidago L., Symphyotrichum Nees
Andrena (Parandrena) andrenoides (Cresson, 1878) IA-IL-IN-KS-KY-MI-MN-MO-ND-NE-OH-OK-SD-TN-TX-WI Uncommon-Rare ...XX....... Salix L.
Andrena (Onagrandrena) anograe Cockerell, 1901 SD Rare ....XXX..... Oenothera L.
Andrena (Cnemidandrena) apacheorum Cockerell, 1897 IL-NE Rare ......XXX... Asteraceae: Cirsium Mill., Ericameria Nutt., Erigeron L., Solidago L.
Andrena (Scaphandrena) arabis Robertson, 1897 AR-IL-IN-KY-MI-MO-OH-TN-WI Common-Uncommon ..XXX....... Arabis L., Cardamine L.
Andrena (Callandrena s.l.) ardis LaBerge, 1967 NE-TX Rare .....XXXXX.. Asteraceae: Chrysothamnus Nutt., Gutierrezia Lag.
Andrena (Parandrena) arenicola LaBerge & Ribble, 1972 KS-OK-TX Rare ...XX....... Salix L.
Andrena (Callandrena s.l.) asteris Robertson, 1891 IA-IL-IN-MI-MN-MO-ND-NE-OH-SD-TN-WI Uncommon .......XX... Solidago L., Symphyotrichum Nees
Andrena (Callandrena s.l.) asteroides Mitchell, 1960 TN Rare ........XXX. Symphyotrichum Nees
Andrena (Euandrena) astragali Viereck & Cockerell, 1914 ND-NE-SD Rare ..XXXX...... Zigadenus Michx.
Andrena (Callandrena s.l.) balsamorhizae LaBerge, 1967 TX Rare ..XX........ Asteraceae: Balsamorhiza Nutt., Enceliopsis (A. Gray) A. Nelson?
Andrena (Callandrena s.l.) beameri LaBerge, 1967 AR-KS-MO Rare .....X...... Asteraceae: Coreopsis L., Echinacea Moench
Andrena (Callandrena s.l.) berkeleyi Viereck & Cockerell, 1914 MO-OK-TX Rare ...XXXXX.... Asteraceae: Engelmannia A. Gray ex Nutt., Hymenoxys Cass., Tetraneuris Greene
Andrena (Thysandrena) bisalicis Viereck, 1908 AR-IL-IN-KY-MI-MN-MO-ND-NE-OH-OK-SD-TN-WI Uncommon-Rare ..XXXXX..... Salix L.
Andrena (Callandrena s.l.) biscutellata Viereck, 1917 KS-OK-TX Rare ...XXX...... Asteraceae: Gaillardia Foug.
Andrena (Conandrena) bradleyi Viereck, 1907 IN-MI-MN-OH-SD-WI Rare ..XXXX...... Ericaceae: Kalmia L., Vaccinium L.
Andrena (Callandrena s.l.) bullata LaBerge, 1967 KS-TX Rare ........XXX. Asteraceae: Heterotheca Cass.
Andrena (Cnemidandrena) canadensis Dalla Torre, 1896 IA-IL-IN-MI-MN-ND-NE-OH-SD-WI Uncommon-Rare .......XX... Asteraceae: Chrysothamnus Nutt., Eurybia (Cass.) Cass., Grindelia Willd., Gutierrezia Lag., Solidago L., Symphyotrichum Nees
Andrena (Scaphandrena) capricornis Casad & Cockerell, 1896 TX Rare ..XX........ Brassicaceae: Descurainia Webb & Bethel., Lepidium L., Lesquerella S. Watson, Physaria (Nutt. ex Torr. & A. Gray) A. Gray
Andrena (Andrena) carolina Viereck, 1909 IN-MI-MN-OH-TN-WI Uncommon ...XXXX..... Ericaceae: Gaylussacia Kunth, Vaccinium L.
Andrena (Callandrena s.l.) chaparralensis Neff & Larkin, 2002 TX Rare .XXX........ Asteraceae: Astereae: Aphanostephus DC., Astranthium Nutt.
Andrena (Conandrena) cheyennorum Viereck & Cockerell, 1914 NE Rare .....XXX.... Symphoricarpos Duham.
Andrena (Cnemidandrena) chromotricha Cockerell, 1899 IL-MI-ND-NE-OH-SD-WI Rare ......XXX... Asteraceae: Grindelia Willd., Helianthus L., Solidago L., Symphyotrichum Nees
Andrena (Andrena) clarkella (Kirby, 1802) MI-MN-ND-WI Rare ...XXXX..... Salix L.
Andrena (Cnemidandrena) colletina Cockerell, 1906 ND-NE Rare .......XX... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Solidago L.
Andrena (Andrena) cornelli Viereck, 1907 TN Rare ....XXX..... Rhododendron L.
Andrena (Cnemidandrena) costillensis Viereck & Cockerell, 1914 SD Rare ......XX.... Asteraceae: Agoseris Raf., Chrysothamnus Nutt., Erigeron L., Grindelia Willd., Heliomeris Nutt., Heterotheca Cass.,
Andrena (Rhacandrena) cragini Cockerell, 1899 KS-IL-MN-MO-ND-NE-SD Rare ....XX...... Amorpha L.
Andrena (Callandrena s.l.) crawfordi Viereck, 1909 IA-KS-ND-OK-TX Uncommon-Rare ..XXX....... Asteraceae: Cichorieae: Pyrrhopappus DC.
Andrena (Dasyandrena) cristata Viereck, 1917 AR-IA-IL-KY Rare .XXXXX...... Arctostaphylos Adans.
Andrena (Diandrena) cuneilabris Viereck, 1926 IL Rare ..XX........ Ranunculus L.
Andrena (Rhaphandrena) dapsilis LaBerge, 1971 OK-TX Rare ...X........ Brassicaceae: Lepidium L., Lesquerella S. Watson, Physaria (Nutt. ex Torr. & A. Gray) A. Gray
Andrena (Ptilandrena) distans Provancher, 1888 IL-IN-MI-MO-NE-OH-TN-WI Uncommon-Rare ...XXX...... Geranium L.
Andrena (Callandrena) duplicata Mitchell, 1960 IL-MO-OH-TN Rare .......XXX.. Asteraceae: Bidens L.
Andrena (Diandrena) eothina Linsley & MacSwain, 1961 IL Rare ..XX........ Camissonia Link?
Andrena (Ptilandrena) erigeniae Robertson, 1891 AR-IA-IL-IN-KY-MI-MO-OH-OK-TN-WI Common ..XXX....... Claytonia L.
Andrena (Tylandrena) erythrogaster (Ashmead, 1890) AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-OK-SD-TN-TX-WI Common-Uncommon ...XXXX..... Salix L.
Andrena (Leucandrena) erythronii Robertson, 1891 IA-IL-IN-KS-MI-MN-MO-NE-OH-TN-WI Uncommon ...XXX...... Erythronium L.
Andrena (Gonandrena) fragilis Smith, 1853 IA-IL-IN-KS-MI-MO-ND-OH-TN-WI Uncommon ....XXX..... Cornus (Swida) L.
Andrena (Andrena) frigida Smith, 1853 IL-IN-MI-MN-ND-NE-SD Common-Uncommon ..XXXX...... Salix L.
Andrena (Callandrena s.l.) fulvipennis Smith, 1853 TX Rare ........XX.. Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
Andrena (Callandrena s.l.) gardineri Cockerell, 1906 AR-IN-KS-MI-MO-SD-TN-TX Uncommon ...XXX...... Gutierrezia Lag., Packera Á. Löve & D. Löve , Senecio L.
Andrena (Euandrena) geranii Robertson, 1891 IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-SD-TN-WI Uncommon ...XXXX..... Hydrophyllum L.
Andrena (Callandrena s.l.) haynesi Viereck & Cockerell, 1914 KS-IA-IL-ND-NE-OK-SD-TX Uncommon-Rare .....XXXX... Helianthus L.
Andrena (Callandrena s.l.) helianthi Robertson, 1891 IA-IL-IN-KS-KY-MI-MN-MO-ND-NE-OH-OK-SD-TX-WI Uncommon-Rare ......XXX... Helianthus L.
Andrena (Callandrena s.l.) helianthiformis Viereck & Cockerell, 1914 KS-MO-ND-NE-SD Rare .....XX..... Echinacea Moench
Andrena (Cnemidandrena) hirticincta Provancher, 1888 IA-IL-IN-MI-MN-ND-NE-OH-SD-WI Uncommon .......XXX.. Asteraceae: Euthamia Nutt. ex. Cass., Grindelia Willd., Solidago L., Symphyotrichum Nees
Andrena (Micrandrena) illinoiensis Robertson, 1891 IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-OK-SD-TX-WI Common-Uncommon ...XXX...... Salix L.
Andrena (Gonandrena) integra Smith, 1853 IA-IL-IN-KS-KY-MI-MN-MO-OH-TN-WI Uncommon-Rare ....XXXXX... Cornus (Swida) L.
Andrena (Callandrena s.l.) irrasus LaBerge, 1967 IL-KS-NE-SD Rare ........X... Asteraceae: Amphiachyris (A. DC.) Nutt., Chrysothamnus Nutt.
Andrena (Micrandrena) ishii Ribble, 1968 KS-IL Rare ..XXX....... Salix L.
Andrena (Tylandrena) jessicae Cockerell, 1896 TX Rare ..XX........ Lesquerella S. Watson
Andrena (Callandrena s.l.) krigiana Robertson, 1901 AR-IL-IN-KY-MI-MN-MO-OH-OK-TN-TX-WI Uncommon ..XXXX...... Cichorieae: Hieracium L., Krigia Schreb.
Andrena (Micrandrena) kristina Lanham, 1983 SD Rare ....X....... Brassicaceae: Lepidium L., Lesquerella S. Watson, Physaria (Nutt. ex Torr. & A. Gray) A. Gray
Andrena (Micrandrena) lamelliterga Ribble, 1968 KY-KS-IL-IN-TN Rare ...XX....... Phacelia Juss.
Andrena (Andrena) macoupinensis Robertson, 1900 AR-IL-IN-KS-MO-NE-ND-OH-OK-SD-TX Uncommon ..XXXX...... Salix L.
Andrena (Trachandrena) mariae Robertson, 1891 AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-SD-TN-WI Uncommon-Rare ..XXX....... Salix L.
Andrena (Micrandrena) melanochroa Cockerell, 1898 AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-OK-SD-TN-TX-WI Uncommon ..XXXXX..... Rosaceae: Fragaria L., Potentilla L., Rubus L.
Andrena (Callandrena s.l.) melliventris Cresson, 1872 KS-ND-OK-TX Rare ...XX....... Asteraceae: Gaillardia Foug.
Andrena (Cnemidandrena) mentzeliae Cockerell, 1897 NE Rare ......XXX... Mentzelia L.
Andrena (Tylandrena) mesillae Cockerell, 1896 OK-TX Rare ..XXXX...... Brassicaceae: Lesquerella S. Watson, Physaria (Nutt. ex Torr. & A. Gray) A. Gray
Andrena (Micrandrena) micheneri Ribble, 1968 TX Rare ..X......... Brassicaceae: Lepidium L., Lesquerella S. Watson, Physaria (Nutt. ex Torr. & A. Gray) A. Gray
Andrena (Callandrena) micheneriella Gusenleitner & Schwartz, 2000 TX Rare ..........X. Asteraceae: Baccharis L.
Andrena (Micrandrena) microchlora Cockerell, 1922 KS-ND-SD Rare ..XXX....... Lomatium Raf.
Andrena (Parandrena) nida Mitchell, 1960 IL-MI-MO-TN Rare ..XXX....... Salix L.
Andrena (Scrapteropsis) nigra Provancher, 1895 OK Rare ..XXXX...... Phacelia Juss.
Andrena (Micrandrena) nigrae Robertson, 1905 AR-IA-IL-IN-KS-KY-MI-MN-MO-ND-NE-OH-OK-SD-TN-TX-WI Uncommon ...XXX...... Salix L.
Andrena (Cnemidandrena) nubecula Smith, 1853 IA-IL-IN-KY-MI-MN-MO-ND-NE-OH-SD-WI Uncommon .......XXX.. Euthamia Nutt. ex Cass., Solidago L., Symphyotrichum Nees
Andrena (Callandrena s.l.) ofella LaBerge, 1967 TX Rare ........XX.. Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Gutierrezia Lag., Haplopappus Cass.
Andrena (Ptilandrena) pallidiscopa Viereck, 1904 IL-NE-SD Rare ..XXXX...... Lomatium Raf.
Andrena (Cnemidandrena) parnassiae Cockerell, 1902 MI-WI Rare .......XX... Parnassia L.
Andrena (Callandrena s.l.) pecosana Cockerell, 1913 TX Rare ......XXX... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Grindelia Willd., Gutierrezia Lag., Haplopappus Cass., Heliomeris Nutt., Heterotheca Cass., Viguiera Kunth, Xanthisma DC.
Andrena (Callandrena) pectidis (Cockerell, 1897) TX Rare ........XX.. Pectis L.
Andrena (Andrena) perarmata Cockerell, 1898 ND Rare .XXXXX...... Salix L.
Andrena (Gonandrena) persimulata Viereck, 1917 IN-MI-MN-ND-NE-WI Uncommon-Rare ....XXX..... Cornus (Swida) L.
Andrena (Euandrena) phaceliae Mitchell, 1960 AR-IL-IN-KS-KY-MO-OH-TN-WI Rare ...XX....... Phacelia Juss.
Andrena (Micrandrena) piperi Viereck, 1904 MO Rare ..XXXX...... Brassicaceae: Descurainia Webb & Bethel., Lepidium L., Lesquerella S. Watson, Physaria (Nutt. ex Torr. & A. Gray) A. Gray
Andrena (Callandrena s.l.) placata Mitchell, 1960 IA-IL-IN-MI-MN-MO-OH-WI Common ......XXXX.. Asteraceae: Astereae: Eurybia (Cass.) Cass., Solidago L., Symphyotrichum Nees
Andrena (Gonandrena) platyparia Robertson, 1895 IA-IL-IN-KS-MI-MN-MO-NE-OH-TX-WI Uncommon-Rare ....XXX..... Cornus (Swida) L.
Andrena (Euandrena) polemonii Robertson, 1891 IA-IL-IN-MI-OH Rare ....X....... Polemonium L.
Andrena (Rhaphandrena) prima Casad, 1896 OK-TX Uncommon-Rare ..XX........ Brassicaceae: Lepidium L., Lesquerella S. Watson, Physaria (Nutt. ex Torr. & A. Gray) A. Gray
Andrena (Micrandrena) primulifrons Casad, 1896 IN-TX Rare ..XXX....... Lesquerella S. Watson
Andrena (Cnemidandrena) ramaleyi Cockerell, 1931 IL Rare .......X.... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt.
Andrena (Callandrena) reflexa Cresson, 1872 TX Rare .......XXX.. Asteraceae: Gutierrezia Lag., Verbesina L.
Andrena (Trachandrena) rehni Viereck, 1907 KS-MI-OH-WI Rare ...XXX...... Castanea Mill.
Andrena (Callandrena s.l.) rudbeckiae Robertson, 1891 AR-IA-IL-IN-KS-KY-MI-MN-MO-ND-NE-OH-OK-TN-TX-WI Rare ....XXXX.... Ratibida Raf., Rudbeckia L..
Andrena (Cnemidandrena) runcinatae Cockerell, 1906 MI-MN-ND Rare .......XX... Asteraceae: Agoseris Raf., Cirsium Mill., Grindelia Willd., Lactuca L., Ratibida Raf.
Andrena (Micrandrena) salictaria Robertson, 1905 IA-IL-IN-KS-KY-MI-MN-MO-ND-NE-OH-SD-WI Uncommon-Rare ..XXX....... Salix L.
Andrena (Scaphandrena) scurra Viereck, 1904 SD Rare ...XXXX..... Brassicaceae: Lepidium L., Lesquerella S. Watson
Andrena (Callandrena s.l.) senticulosa LaBerge, 1967 TX Rare ..XX........ Asteraceae: Berlandiera DC., Engelmannia A. Gray ex Nutt., Pyrrhopappus DC., Krigia Schreb.
Andrena (Trachandrena) sigmundi Cockerell, 1902 AR-IA-IL-IN-MI-MN-ND-NE-OH-SD-WI Uncommon ..XXXX...... Salix L.
Andrena (Callandrena s.l.) simplex Smith, 1853 AR-IA-IL-IN-KS-KY-MI-MN-MO-NE-OH-OK-TN-WI Common ......XXXXX. Solidago L., Symphyotrichum Nees
Andrena (Callandrena) simulata Smith, 1879 TX Rare .......XX... Asteraceae: Viguiera Kunth
Andrena (Callandrena s.l.) sitiliae Viereck, 1909 OK-TX Rare ...XX....... Asteraceae: Pyrrhopappus DC.
Andrena (Cnemidandrena) sulcata Donovan, 1977 ND-WI Rare .......XX... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt.
Andrena (Cnemidandrena) surda Cockerell, 1910 ND-NE-SD Uncommon-Rare .....XXXX... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Gutierrezia Lag., Machaeranthera Nees, Solidago L.
Andrena (Callandrena s.l.) texana Cresson, 1872 TX Rare .........X.. Asteraceae: Amphiachyris (A. DC.) Nutt.
Andrena (Callandrena s.l.) tonkaworum Viereck, 1917 KS-NE-OK-TX Rare ...XXXX..... Asteraceae: Engelmannia A. Gray ex Nutt.
Andrena (Micrandrena) trapezoidea Viereck, 1917 KS-OK-TX Rare ..XX........ Lesquerella S. Watson
Andrena (Callandrena) trimaculata LaBerge, 1967 TX Rare .......XXX.. Asteraceae: Helianthus L.
Andrena (Thysandrena) trizonata Ashmead, 1890 ND Rare ....XXX..... Salix L.
Andrena (Callandrena) verbesinae Viereck & Cockerell, 1914 TX Rare .......XXXX. Asteraceae: Grindelia Willd., Heliomeris Nutt., Verbesina L.
Andrena (Callandrena s.l.) verecunda Cresson, 1872 OK-TX Rare ...X........ Asteraceae: Engelmannia A. Gray ex Nutt., Gaillardia Foug. Pyrrhopappus DC.
Andrena (Iomelissa) violae Robertson, 1891 AR-IA-IL-IN-KS-KY-MI-MO-NE-OH-SD-TN-WI Common-Uncommon ..XX........ Viola L.
Andrena (Callandrena s.l.) vulpicolor Cockerell, 1897 SD-TX Rare ......XXXX.. Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Heliomeris Nutt., Heterotheca Cass., Senecio L., Viguiera Kunth
Andrena (Parandrena) wellesleyana Robertson, 1897 IL-IN-MI-MN-ND-NE-WI Uncommon-Rare ...XX....... Salix L.
Andrena (Micrandrena) ziziae Robertson, 1891 IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-OK-SD-TN-WI Uncommon ...XXX...... Zizia W.D.J. Koch [Taenidia (Torr. & A. Gray) Drude, Thaspium Nutt.?]
Andrena (Derandrena) ziziaeformis Cockerell, 1908 IN-MO-OH-TN Uncommon-Rare ...XX....... Potentilla L., Waldsteinia Willd.
Andrenidae: Panurginae: Calliopsini Calliopsis (Hypomacrotera) callops (Cockerell & Porter, 1899) KS-TX Uncommon-Rare ...XXXXXX... Chamaesaracha (A. Gray) Benth., Physalis L.
Calliopsis (Nomadopsis) callosa (Timberlake, 1952) TX Rare .......X.... Tidestromia Standl.
Calliopsis (Calliopsima) coloradensis Cresson, 1878 AR-IA-IL-KS-MN-MO-ND-NE-SD-TX-WI Uncommon-Rare .....XXXXX.. Asteraceae: Chrysopsis (Nutt.) Elliot, Chrysothamnus Nutt., Ericameria Nutt., Grindelia Willd., Gutierrezia Lag., Helianthus L., Heliomeris Nutt., Heterotheca Cass., Xanthisma DC.
Calliopsis (Calliopsima) coloratipes Cockerell, 1898 TX Rare .......XXX.. Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Haplopappus Cass., Helianthus L., Heterotheca Cass., Verbesina L., Viguiera Kunth
Calliopsis (Calliopsima) crypta Shinn, 1965 TX Rare ......XX.... Asteraceae: Cirsium Mill., Heterotheca Cass., Verbesina L.
Calliopsis (Nomadopsis) helianthi (Swenk & Cockerell, 1907) KS-NE Rare ...XXXXXX... Euphorbiaceae: Chamaesyce Gray, Euphorbia L.
Calliopsis (Verbenapis) hirsutifrons Cockerell, 1896 SD-TX Rare .....XX..... Verbena L.
Calliopsis (Nomadopsis) meliloti Cockerell, 1896 TX Uncommon-Rare ....XXXXX... Fabaceae: Dalea L., Psorothamnus Rydb.
Calliopsis (Verbenapis) michenerella Shinn & Engel, 2003 TX Rare ..XX........ Verbena L.
Calliopsis (Nomadopsis) micheneri (Rozen, 1958) TX Rare ...XXXX..... Trifolium L.
Calliopsis (Verbenapis) nebraskensis Crawford, 1902 AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-SD-WI Uncommon-Rare ......XX.... Verbena L.
Calliopsis (Calliopsima) philiphunteri Shinn & Engel, 2003 TX Rare .......XXX.. Asteraceae: Ericameria Nutt., Grindelia Willd., Gutierrezia Lag., Isocoma Nutt., Machaeranthera Nees, Verbesina L.
Calliopsis (Nomadopsis) puellae (Cockerell, 1933) TX Rare ..XXXX...... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Calycoseris A. Gray, Chaenactis DC., Malacothrix DC.
Calliopsis (Calliopsima) rozeni Shinn, 1965 TX Uncommon .......XXX.. Asteraceae: Bahia Lag., Baileya Harv. & A. Gray ex A. Gray, Dyssodia Cav., Encelia Adans., Gutierrezia Lag., Heliomeris Nutt., Heterotheca Cass., Machaeranthera Nees, Malacothrix DC., Psilostrophe DC., Verbesina L.
Calliopsis (Hypomacrotera) subalpina Cockerell, 1894 KS-OK-TX Common ..XXXXXXX... Sphaeralcea A. St.-Hil.
Calliopsis (Verbenapis) verbenae Cockerell & Porter, 1899 KS-TX Rare ....XXXXX... Verbena L.
Calliopsis (Nomadopsis) zebrata Cresson, 1878 OK-MN-SD Rare .....XXX.... Astragalus L.
Calliopsis (Nomadopsis) zonalis Cresson, 1879 KS Rare .....XX..... Monardella Benth.
Andrenidae: Panurginae: Panurgini: Panurginia Panurginus atramontensis Crawford, 1926 MO-TN Rare ...XXX...... Vaccinium (Polycodium) L.
Panurginus nigrihirtus Michener, 1936 KS Rare ..XX........ Ranunculus L.
Panurginus potentillae (Crawford, 1916) IL-KY-MO-NE-TN Uncommon-Rare ..XXX....... Fragaria L., Potentilla L.
Andrenidae: Panurginae: Panurgini: Perditina Macrotera (Macroteropsis) haplura (Cockerell, 1922) TX Rare ........X... Sphaeralcea A. St.-Hil.
Macrotera (Macroteropsis) latior (Cockerell, 1896) TX Rare .....XXXX... Malvaceae: Sphaeralcea A. St.-Hil.
Macrotera (Macroteropsis) magniceps (Timberlake, 1960) TX Rare .......X.... Malvaceae: Sphaeralcea A. St.-Hil.
Macrotera (Macroterella) mellea (Timberlake, 1954) TX Rare ......XXX... Euphorbiaceae: Chamaesyce Gray, Euphorbia L.
Macrotera (Cockerellula) opuntiae (Cockerell, 1922) ND-SD-TX Rare .....X...... Opuntia Mill.
Macrotera (Macroteropsis) portalis (Timberlake, 1954) TX Rare ....XXXXX... Sphaeralcea A. St.-Hil.
Macrotera (Macrotera) texana Cresson, 1878 OK-TX Rare ....XX...... Opuntia Mill.
Andrenidae: Panurginae: Panurgini: Perditina Perdita (Perdita) affinis Cresson, 1878 TX Rare ......XX.... Asteraceae: Chrysothamnus Nutt., Grindelia Willd., Senecio L., Xanthisma DC.
Perdita (Cockerellia) albihirta Timberlake, 1954 TX Rare ..XXX....... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chaenactis DC., Encelia Adans.
Perdita (Cockerellia) albipennis Cresson, 1868 IL-IN-KS-MI-MN-MO-ND-NE-OK-SD-TX-WI Uncommon-Rare .....XXXXX.. Asteraceae: Grindelia Willd., Helianthus L., Heterotheca Cass., Rudbeckia L., Symphyotrichum Nees, Verbesina L.
Perdita (Pentaperdita) albovittata Cockerell, 1895 TX Uncommon ......XXX... Asteraceae: Bahia Lag., Baileya Harv. & A. Gray ex A. Gray, Gutierrezia Lag., Heterotheca Cass., Hymenothrix A. Gray, Isocoma Nutt., Machaeranthera Nees, Pectis L., Psilostrophe DC., Verbesina L., Xanthisma DC.
Perdita (Hexaperidita) alexi Timberlake, 1968 KS-OK-TX Rare .........X.. Asteraceae: Helianthus L., Heterotheca Cass.
Perdita (Perdita) ampla Timberlake, 1971 TX Rare ........X... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt.
Perdita (Perdita) aperta Timberlake, 1968 TX Rare ........XX.. Gutierrezia Lag.
Perdita (Perdita) ashmeadi Cockerell, 1899 TX Uncommon ..XXX....... Prosopis L.
Perdita (Hexaperdita) asteris Cockerell, 1896 TX Uncommon-Rare .....XXXXX.. Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Dieteria Nutt., Heterotheca Cass.
Perdita (Cockerellia) beata Cockerell, 1895 TX Rare .....XXXX... Asteraceae: Bahia Lag., Encelia Adans., Gutierrezia Lag., Verbesina L.
Perdita (Cockerellia) bequaerti Viereck, 1917 AR-IL-IN-MI-MN-MO-OK-TN-TX-WI Rare .....XXXX... Asteraceae: Bidens L., Balduina Nutt., Helianthus L.[Physalis L.?]
Perdita (Xerophasma) bequaertiana Cockerell, 1951 TX Rare .....XXX.... Oenothera L.
Perdita (Hexaperdita) bishoppi Cockerell, 1906 KS-NE-OK-TX Rare ......XXX... Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
Perdita (Hexaperdita) boltoniae (Robertson, 1902) IL Rare ......XXXX.. Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
Perdita (Perdita) bruneri Cockerell, 1897 KS-ND-NE-TX Uncommon-Rare .......XX... Asteraceae: Grindelia Willd., Helianthus L., Heterotheca Cass., Solidago L., Symphyotrichum Nees
Perdita (Hexaperdita) callicerata Cockerell, 1897 TX Common-Uncommon ....XXXXX... Asteraceae: Bahia Lag., Baileya Harv. & A. Gray ex A. Gray, Gutierrezia Lag., Heterotheca Cass., Malacothrix DC., Verbesina L.
Perdita (Hexaperdita) cambarella Cockerell, 1906 KS-OK-TX Rare ......XX.... Asteraceae: Croptilon Raf., Heterotheca Cass.
Perdita (Pentaperdita) chrysophila Cockerell, 1896 TX Rare ........X... Asteraceae: Thelesperma Less., Verbesina L.
Perdita (Perditella) cladothricis Cockerell, 1896 OK-TX Common-Uncommon ......XXX... Tidestromia Standl.
Perdita (Perdita) confusa Timberlake, 1964 TX Uncommon-Rare ....XXXX.... Lepidium L.
Perdita (Cockerellia) coreopsidis Cockerell, 1906 KS-NE-OK-TX Uncommon-Rare ...XXXXX.... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Gaillardia Foug., Heterotheca Cass.
Perdita (Perdita) crotonis Cockerell, 1896 KS-NE-TX Uncommon-Rare .....XXXX... Croton L.
Perdita (Perdita) dalyi Timberlake, 1960 TX Rare .......X.... Helianthus L.
Perdita (Perdita) difficilis Timberlake, 1964 TX Uncommon ..XXXX...... Prosopis L.
Perdita (Epimacrotera) dolanensis Neff, 2003 TX Rare ...X........ Salvia L.
Perdita (Perdita) dolichocephala Swenk & Cockerell, 1907 KS-NE Uncommon-Rare ........X... Asteraceae: Helianthus L.
Perdita (Perdita) exclamans Cockerell, 1895 TX Rare ..XXX....... Fabaceae: Prosopis L.
Perdita (Perdita) fallax Cockerell, 1896 NE-TX Rare ......XXX... Asteraceae: Bahia Lag., Chrysothamnus Nutt., Erigeron L., Gaillardia Foug., Grindelia Willd., Gutierrezia Lag., Haplopappus Cass., Helianthus L., Heterotheca Cass., Solidago L.
Perdita (Hexaperdita) foveata Timberlake, 1956 AR-KS-OK-TX Rare ....XX...... Asteraceae: Coreopsis L.
Perdita (Perdita) georgica Timberlake, 1928 IL Uncommon .......XXX.. Chrysopsis (Nutt.) Elliot, Heterotheca Cass., Pityopsis Nutt.
Perdita (Perdita) gerhardi Viereck, 1904 IL-IN-MI-TX-WI Uncommon-Rare ...XXXXX.... Monarda L.
Perdita (Perdita) gutierreziae Cockerell, 1896 TX Uncommon-Rare .....XXXXX.. Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Ericameria Nutt., Grindelia Willd., Gutierrezia Lag., Isocoma Nutt., Machaeranthera Nees, Sartwellia A. Gray, Solidago L.
Perdita (Perdita) halictoides Smith, 1853 AR-IA-IL-IN-KS-MI-MN-ND-NE-OK-SD-TX Uncommon-Rare ...XXXXXX... Physalis L.
Perdita (Perdita) heliotropii Cockerell, 1900 TX Rare ...XXX...... Heliotropium L.
Perdita (Heteroperdita) hippolyta Portman & Griswold, 2016 TX Rare ....XXXXX... Tiquilia Pers.
Perdita (Heteroperdita) hooki Portman & Neff, 2016 TX Rare ......XXX... Tiquilia Pers.
Perdita (Hexaperdita) ignota Cockerell, 1896 AR-KS-MO-NE-OK-TX Uncommon-Rare .....XXX.... Asteraceae: Astereae: Grindelia Willd., Heterotheca Cass., Solidago L.
Perdita (Perdita) kiowi Griswold, 1988 KS-NE-TX Rare .....XXX.... Mentzelia L.
Perdita (Perdita) labergei Timberlake, 1960 KS Rare ......XX.... Euphorbia L.
Perdita (Perditella) larreae Cockerell, 1896 TX Uncommon-Rare ......XXX... Larrea Cav.
Perdita (Perdita) lasiogastra Timberlake, 1929 TX Rare .......XX... Pectis L.
Perdita (Perdita) laticincta Swenk & Cockerell, 1907 KS-ND-NE-SD Uncommon-Rare ......XX.... Asteraceae: Helianthus L., Solidago L.
Perdita (Perdita) lenis Timberlake, 1958 TX Rare ..XXXXXX.... Solanaceae: Chamaesaracha (A. Gray) Benth., Physalis L., Solanum L.
Perdita (Cockerellia) lepachidis Cockerell, 1896 KS-NE-OK-TX Rare ....XXXX.... Asteraceae: Ratibida Raf.
Perdita (Cockerellia) lingualis Cockerell, 1896 KS-ND-NE-OK-SD-TX Uncommon-Rare ......XX.... Asteraceae: Ericameria Nutt., Helianthus L., Verbesina L.
Perdita (Perdita) luciae Cockerell, 1899 TX Rare ...XXX...... Fabaceae: Parkinsonia L., Prosopis L.
Perdita (Perdita) luteola Cockerell, 1894 NE-TX Rare .......XX... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Ericameria Nutt., Gutierrezia Lag., Isocoma Nutt., Xanthocephalum Willd.
Perdita (Perdita) maculigera Cockerell, 1896 IA-IL-IN-KS-MI-MN-MO-OH-NE-SD-TX-WI Uncommon-Rare ...XXX...... Salix L.
Perdita (Pygoperdita) malacothricis Timberlake, 1956 TX Rare ..XXX....... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chaenactis DC., Malacothrix DC., Rafinesquia Nutt., Stephanomeria Nutt.
Perdita (Perdita) melanostoma Swenk & Cockerell, 1907 NE Rare ......XX... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Ericameria Nutt., Gutierrezia Lag.
Perdita (Perdita) mentzeliarum Cockerell, 1897 TX Rare ....XXXXX... Mentzelia L.
Perdita (Perditella) minima Cockerell, 1923 TX Uncommon-Rare ......XXX... Euphorbiaceae: Chamaesyce Gray, Euphorbia L.
Perdita (Perdita) munita Timberlake, 1964 TX Rare .......X.... Chamaesaracha (A. Gray) Benth.
Perdita (Perdita) nasuta Timberlake, 1962 TX Uncommon-Rare .....XXX.... Eriogonum Michx.
Perdita (Hexaperdita) nubila Timberlake, 1958 AR-KS-TN Rare ....X....... Asteraceae: Erigeron L.
Perdita (Perdita) numerata Cockerell, 1895 TX Rare ..XX........ Fabaceae: Prosopis L.
Perdita (Alloperdita) obscurata Cresson, 1878 IL-MO-TX Rare ....X....... Diospyros L., Vaccinium L.
Perdita (Perdita) octomaculata (Say, 1824) AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-OK-WI Uncommon-Rare ......XXXX.. Asteraceae: Astereae: Baccharis L., Chrysopsis (Nutt.) Elliot, Euthamia Nutt. ex Cass., Grindelia Willd., Pityopsis Nutt., Solidago L., Symphyotrichum Nees
Perdita (Epimacrotera) peculiaris Timberlake, 1953 TX Rare ...XXXXXX... Chamaesaracha (A. Gray) Benth.
Perdita (Perdita) pedernalensis Neff, 2010 TX Rare ...X........ Rubiaceae: Cephalanthus L., Diodia L., Galium L., Houstonia L., Mitchella L.
Perdita (Perdita) perpallida Cockerell, 1901 KS-MN-ND-NE-OK-WI Rare .....XX..... Dalea L.
Perdita (Cockerellia) perpulchra Cockerell, 1896 KS-TX Uncommon-Rare .....XXXX... Asteraceae: Aphanostephus DC., Baileya Harv. & A. Gray ex A. Gray, Dyssodia Cav., Grindelia Willd., Gutierrezia Lag., Heterotheca Cass., Isocoma Nutt., Verbesina L., Xanthocephalum Willd., Xylothamia G.L. Nesom, Suh, D. Morgan & Simpson
Perdita (Perdita) phymatae Cockerell, 1895 KS Uncommon-Rare .......XX... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Ericameria Nutt., Gutierrezia Lag., Helianthus L.
Perdita (Heteroperdita) pilonotata Timberlake, 1980 TX Rare .....XXXX... Tiquilia Pers.
Perdita (Hexaperdita) pratti Cockerell, 1906 KS-TX Uncommon .....XXXXXXX Asteraceae: Helianthus L., Heterotheca Cass.
Perdita (Perdita) prionopsidis Timberlake, 1960 KS Rare ........X... Asteraceae: Boltonia L'Hér., Grindelia Willd., Heterotheca Cass.
Perdita (Perdita) punctosignata Cockerell, 1895 TX Uncommon ..XXXX...... Fabaceae: Parkinsonia L., Prosopis L.
Perdita (Cockerellia) purpurascens Timberlake, 1956 TX Uncommon-Rare ....XX...... Asteraceae: Gaillardia Foug.
Perdita (Perdita) rhodura Cockerell, 1897 NE-TX Uncommon-Rare .......XX... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Ericameria Nutt., Gutierrezia Lag., Viguiera Kunth
Perdita (Perdita) semicaerulea Cockerell, 1896 TX Uncommon ...XXXXXX... Larrea Cav.
Perdita (Cockerellia) shinnersi Timberlake, 1956 TX Rare ....X....... Asteraceae: Gaillardia Foug.
Perdita (Perdita) sphaeralceae Cockerell, 1896 TX Uncommon .....XXXX... Sphaeralcea A. St.-Hil.
Perdita (Perdita) stathamae Timberlake, 1964 TX Uncommon-Rare ...XXX...... Fabaceae: Prosopis L.
Perdita (Perdita) swenki Crawford, 1915 IL-IN-MI-MN-ND-NE-WI Uncommon-Rare ......XXX... Asteraceae: Chrysopsis (Nutt.) Elliot, Grindelia Willd., Helianthus L., Heterotheca Cass., Solidago L.
Perdita (Perdita) taeniata Timberlake, 1971 TX Rare ........X... Ericameria Nutt.
Perdita (Perdita) translineata Timberlake, 1960 NE-TX Rare ....XXXXX... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Heterotheca Cass.
Perdita (Perdita) triangulifera Timberlake, 1964 TX Uncommon-Rare ..XXX....... Fabaceae: Parkinsonia L., Prosopis L.
Perdita (Perdita) tridentata Stevens, 1919 IA-KS-ND-NE Rare .......XX... Helianthus L.
Perdita (Heteroperdita) trifasciata Timberlake, 1953 TX Rare ...XXXXXXXXX Tiquilia Pers.
Perdita (Perdita) trinotata Timberlake, 1964 TX Rare ...XX....... Lesquerella S. Watson
Perdita (Perdita) variegata Timberlake, 1960 KS-NE-TX Uncommon-Rare .....XXX.... Monarda L.
Perdita (Cockerellia) verbesinae Cockerell, 1896 TX Uncommon-Rare ......XXX... Asteraceae: Baccharis L., Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Gutierrezia Lag., Helianthus L., Heliopsis Pers., Heterotheca Cass., Verbesina L., Viguiera Kunth
Perdita (Perdita) wislizeniae Timberlake, 1964 TX Rare .......XX... Capparaceae: Cleome L., Cleomella DC., Wislizenia Engelm.
Perdita (Perdita) wootonae Cockerell, 1898 KS-NE-TX Rare .......XX... Mentzelia L.
Perdita (Hexaperdita) xanthismae Cockerell, 1905 KS-NE-OK-SD-TX Rare ........X... Asteraceae: Xanthisma DC.
Perdita (Perdita) zebrata Cresson, 1878 ND-NE Uncommon ......XX.... Capparaceae: Cleome L., Cleomella DC., Wislizenia Engelm.
Perdita (Perdita) zonalis Cresson, 1879 SD Uncommon-Rare .......XX... Asteraceae: Chrysothamnus Nutt., Dieteria Nutt., Ericameria Nutt., Eriogonum Michx., Gutierrezia Lag., Heterotheca Cass., Isocoma Nutt., Lepidospartum (A. Gray) A. Gray, Solidago L., Symphyotrichum Nees
Andrenidae: Panurginae: Protandrenini Protandrena abdominalis (Cresson, 1878) KS-OK-TX Uncommon-Rare ....XXXXX... Monarda L.
Protandrena verbesinae (Timberlake, 1955) TX Rare .......X.... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Helianthus L., Heterotheca Cass., Machaeranthera Nees, Pectis L.
Andrenidae: Panurginae: Protandrenini Pseudopanurgus aestivalis (Provancher, 1882) IL-IN-MI-MN-ND-NE-SD-WI Rare .......XX... Asteraceae: Astereae: Chrysopsis (Nutt.) Elliot, Erigeron L., Grindelia Willd., Solidago L., Symphyotrichum Nees
Pseudopanurgus aethiops (Cresson, 1872) KS-NE-OK-TX Uncommon-Rare .......XX... Asteraceae: Helianthus L., Heterotheca Cass., Verbesina L., Viguiera Kunth
Pseudopanurgus albitarsis (Cresson, 1872) IA-IL-KS-MI-ND-OK-SD-TX-WI Rare ....XXXXX... Asteraceae: Baccharis L., Coreopsis L., Echinacea Moench, Helianthus L., Ratibida Raf., Rudbeckia L..
Pseudopanurgus andrenoides (Smith, 1853) IL-IN-KS-MI-MN-MO-ND-OH-WI Uncommon .......XX... Asteraceae: Eurybia (Cass.) Cass., Rudbeckia L., Solidago L., Symphyotrichum Nees
Pseudopanurgus boylei (Cockerell, 1896) TX Rare .......XXX.. Asteraceae: Verbesina L.
Pseudopanurgus compositarum (Robertson, 1893) AR-IA-IL-IN-KS-KY-MI-MO-OH-OK-TN-TX Uncommon ........XX.. Asteraceae: Bidens L., Rudbeckia L., Solidago L., Symphyotrichum Nees
Pseudopanurgus fraterculus (Cockerell, 1896) TX Rare .......XX... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Gutierrezia Lag., Haplopappus Cass., Heliopsis Pers., Heterotheca Cass., Hymenothrix A. Gray, Machaeranthera Nees, Melampodium L., Verbesina L.
Pseudopanurgus helianthi Mitchell, 1960 IN Rare .......X.... Asteraceae: Helianthus L.
Pseudopanurgus illinoiensis (Cresson, 1878) AR-IL-IN-KS-OK-TN-TX Rare ....X....... Asteraceae: Erigeron L., Heterotheca Cass.
Pseudopanurgus illustris Timberlake, 1967 KS-MO-ND-NE-SD Rare .......XX... Asteraceae: Verbesina L.
Pseudopanurgus innuptus (Cockerell, 1896) KS-MO-ND-NE-SD Rare .......X.... Asteraceae: Helianthus L.
Pseudopanurgus labrosiformis (Robertson, 1898) KS-IL-MI-MO-NE Rare .......XXX.. Asteraceae: Bidens L., Coreopsis L., Helianthus L., Rudbeckia L., Silphium L., Solidago L.
Pseudopanurgus labrosus (Robertson, 1895) IL-IN-MN-MO-ND-OH-WI Rare .......XX... Asteraceae: Helianthus L., Rudbeckia L..
Pseudopanurgus leucopterus (Cockerell, 1923) NE Rare .......X.... Asteraceae: Helianthus L.
Pseudopanurgus occiduus Timberlake, 1967 TX Rare .......XX... Asteraceae: Heterotheca Cass.
Pseudopanurgus passiflorae Robertson, 1897 IL-MO-OH-TX Rare ...XXXXXX... Passiflora L.
Pseudopanurgus pauper (Cresson, 1878) IL-IN-MN-MO-TN Rare .....XXX.... Ceanothus L.
Pseudopanurgus piercei (Crawford, 1903) ND-NE Rare .......XX... Asteraceae: Grindelia Willd., Helianthus L., Solidago L.
Pseudopanurgus renimaculatus (Cockerell, 1896) KS-ND-NE-SD-TX Rare .......XXX.. Asteraceae: Baccharis L., Boltonia L'Hér., Grindelia Willd., Helianthus L., Symphyotrichum Nees
Pseudopanurgus rugosus (Robertson, 1895) AR-IL-IN-KS-MI-MO-OK-TN-TX Rare .......XX... Asteraceae: Bidens L., Helianthus L., Heliopsis Pers., Rudbeckia L., Silphium L.
Pseudopanurgus simulans (Swenk & Cockerell, 1907) MN-ND-NE Rare ........X... Asteraceae: Grindelia Willd., Helianthus L.
Pseudopanurgus solidaginis (Robertson, 1893) IL Rare .......XXX.. Asteraceae: Bidens L., Coreopsis L., Helianthus L., Pityopsis Nutt., Rudbeckia L., Solidago L.
Pseudopanurgus virginicus (Cockerell, 1907) IN Rare ....XX...... Ceanothus L., Houstonia L.
Apidae: Apinae: Anthophorini Anthophora (Lophanthophora) affabilis Cresson, 1878 KS-SD-TX Uncommon-Rare ...XXX...... Oenothera L.
Anthophora (Micranthophora) curta Provancher, 1895 TX Uncommon-Rare ....XXXX.... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Bebbia (Benth.) Greene, Coreopsis L., Encelia Adans., Ericameria Nutt., Gaillardia Foug., Grindelia Willd., Gutierrezia Lag., Haplopappus Cass., Helianthus L., Heterotheca Cass., Hymenothrix A. Gray, Machaeranthera Nees, Palafoxia Lag., Solidago L., Stephanomeria Nutt., Verbesina L.
Anthophora (Lophanthophora) porterae Cockerell, 1900 KS-OK-SD-TX Uncommon-Rare ..XXXX...... Astragalus L.
Apidae: Apinae: Anthophorini Habropoda laboriosa (Fabricius, 1804) AR-IL-ID-KS-MI-OH-OK-TN-TX-WI Uncommon-Rare XXXXX.....XX Cercis L., Vaccinium L.
Apidae: Apinae: Emphorini Ancyloscelis apiformis (Fabricius, 1793) TX Uncommon-Rare ..XXXXX..... Ipomoea L.
Ancyloscelis sejunctus Cockerell, 1933 TX Rare .......XX... Ipomoea L.
Apidae: Apinae: Emphorini Diadasia (Coquillettapis) afflicta (Cresson, 1878) KS-NE-OK-TX Rare ..XXXX...... Malvaceae: Callirhoe Nutt.
Diadasia (Coquillettapis) afflictula Cockerell, 1910 TX Rare .......XX... Malvaceae: Malvella Jaubert & Spach, Sida L.
Diadasia (Coquillettapis) australis (Cresson, 1878) KS-ND-NE-SD-TX Common-Uncommon ...XXXXXXXX. Cactaceae: Cylindropuntia (Engelm.) Kreuzinger, Echinocactus Link & Otto, Ferocactus Britton & Rose, Opuntia Mill.
Diadasia (Coquillettapis) diminuta (Cresson, 1878) KS-ND-NE-OK-SD-TX Common .....XXX.... Malvaceae: Sidalcea A. Gray, Sphaeralcea A. St.-Hil.
Diadasia (Diadasia) enavata (Cresson, 1872) AR-IL-KS-MO-ND-NE-OK-SD-TX Uncommon-Rare ......XXXX.. Asteraceae: Helianthus L.
Diadasia (Coquillettapis) lutzi Cockerell, 1924 TX Uncommon ...XXX...... Malvaceae: Sphaeralcea A. St.-Hil.
Diadasia (Coquillettapis) megamorpha Cockerell, 1898 TX Rare .......XX... Sphaeralcea A. St.-Hil.
Diadasia (Dasiapis) ochracea (Cockerell, 1903) TX Common-Uncommon ....XXXXX... Malvaceae: Malacothamnus Greene, Malva L., Malvella Jaubert & Spach, Sphaeralcea A. St.-Hil.
Diadasia (Coquillettapis) piercei Cockerell, 1911 TX Rare ..XXXX...... Opuntia Mill.
Diadasia (Coquillettapis) rinconis Cockerell, 1897 TX Common-Uncommon ..XXXXXX.... Cactaceae: Cylindropuntia (Engelm.) Kreuzinger, Echinocactus Link & Otto, Ferocactus Britton & Rose, Opuntia Mill.
Diadasia (Coquillettapis) sphaeralcearum Cockerell, 1905 NE-TX Uncommon-Rare ..XXXXXXXXX. Sphaeralcea A. St.-Hil.
Diadasia (Coquillettapis) vallicola Timberlake, 1940 TX Uncommon-Rare ..XXX....... Sphaeralcea A. St.-Hil.
Apidae: Apinae: Emphorini Melitoma grisella (Cockerell & Porter, 1899) KS-ND-NE-OK-SD-TX Rare .....XXX.... Ipomoea L.
Melitoma marginella (Cresson, 1872) TX Rare ...XXXX..... Ipomoea L.
Melitoma taurea (Say, 1837) IA-IL-IN-KS-KY-MO-OH-TN-TX Uncommon .....XXXX... Calystegia R. Br., Ipomoea L.
Apidae: Apinae: Emphorini Ptilothrix bombiformis (Cresson, 1878) AR-IA-IL-IN-KS-KY-MO-OH-TN-TX Common-Uncommon .....XXX.... Hibiscus L.
Apidae: Apinae: Eucerini Cemolobus ipomoeae (Robertson, 1891) IL-IN-MO-OH Rare .....XX..... Calystegia R. Br., Ipomoea L.
Apidae: Apinae: Eucerini Florilegus (Florilegus) condignus (Cresson, 1878) IL-IN-KS-MI-MO-NE-OK-TX-WI Uncommon-Rare ...XXXXX.... Pontederia L.
Apidae: Apinae: Eucerini Melissodes (Eumelissodes) agilis Cresson, 1878 AR-IA-IL-IN-KS-KY-MI-MN-MO-ND-NE-OH-OK-SD-TN-TX-WI Common-Uncommon ....XXXXXXX.Asteraceae: Cirsium Mill., Grindelia Willd., Helianthus L., Rudbeckia L., Ratibida Raf., Simsia Pers., Solidago L., Verbesina L.
Melissodes (Apomelissodes) apicatus Lovell & Cockerell, 1906 OH Rare ..XXXXX..... Pontederia L.
Melissodes (Eumelissodes) bidentis Cockerell, 1914 IA-IL-MN-MO-ND-NE-SD-TX Rare ......XXX... Asteraceae: Bidens L., Helianthus L., Rudbeckia L..
Melissodes (Eumelissodes) bimatris LaBerge, 1961 KS Rare .......XXX..Asteraceae: Chrysothamnus Nutt., Cirsium Mill., Dieteria Nutt., Ericameria Nutt., Grindelia Willd., Gutierrezia Lag., Helianthus L., Isocoma Nutt., Symphyotrichum Nees, Verbesina L.
Melissodes (Eumelissodes) boltoniae Robertson, 1905 AR-IA-IL-IN-KS-KY-MN-NE-TX Uncommon ......XXXXX. Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Cirsium Mill., Coreopsis L., Grindelia Willd., Helianthus L., Heterotheca Cass., Ratibida Raf., Rudbeckia L., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
Melissodes (Callimelissodes) coloradensis Cresson, 1878 IL-IN-KS-MO-NE-OH-SD-WI Rare .......XX...Asteraceae: Cirsium Mill., Coreopsis L., Helianthus L., Heliopsis Pers., Liatris Gaertn. ex Schreb., Machaeranthera Nees, Rudbeckia L., Silphium L., Solidago L., Vernonia Schreb.
Melissodes (Eumelissodes) confusus Cresson, 1878 MN-ND-TX Rare .......XX...Asteraceae: Bidens L., Chrysothamnus Nutt., Cirsium Mill., Cosmos Cav., Erigeron L., Grindelia Willd., Gutierrezia Lag., Helianthus L., Heliomeris Nutt., Heliopsis Pers., Heterotheca Cass., Hymenoxys Cass., Pericome A. Gray, Pyrrocoma Hook., Rudbeckia L., Senecio L., Simsia Pers., Symphyotrichum Nees
Melissodes (Eumelissodes) coreopsis Robertson, 1905 AR-IA-IL-KS-MN-MO-ND-NE-OH-OK-SD-TX-WI Common ......XXXX..Asteraceae: Cirsium Mill., Coreopsis L., Echinacea Moench, Erigeron L., Gaillardia Foug., Grindelia Willd., Gutierrezia Lag., Helianthus L., Heterotheca Cass., Machaeranthera Nees, Ratibida Raf., Solidago L., Symphyotrichum Nees, Verbesina L.
Melissodes (Eumelissodes) denticulatus Smith, 1854 AR-IA-IL-IN-KS-MO-ND-NE-OH-TN Common-Uncommon ....XXXXXX.. Asteraceae: Vernonia Schreb.
Melissodes (Eumelissodes) dentiventris Smith, 1854 AR-IL-IN-KS-KY-MI-MO-ND-OH-OK-TN-TX-WI Uncommon ......XXXX.. Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Coreopsis L., Helianthus L., Pityopsis Nutt., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
Melissodes (Heliomelissodes) desponsus Smith, 1854 AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-WI Uncommon .....XXXXX.. Asteraceae: Cirsium Mill.
Melissodes (Eumelissodes) druriellus (Kirby, 1802) IA-IL-MI-MN-OH-ND-TN-WI Uncommon ....XXXXXXX. Asteraceae: Euthamia Nutt. ex Cass., Helianthus L., Rudbeckia L., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
Melissodes (Apomelissodes) fimbriatus Cresson, 1878 KS-TX Rare ...XXXX..... Oenothera L.
Melissodes (Eumelissodes) fumosus LaBerge, 1961 IL-IN-KS-NE-TX Rare .....XXXXX.. Asteraceae: Chrysopsis (Nutt.) Elliot, Pityopsis Nutt., Solidago L.
Melissodes (Eumelissodes) gelidus LaBerge, 1961 IL-KS-ND-NE-SD-TX Rare .....XX..... Asteraceae: Helianthus L.
Melissodes (Callimelissodes) glenwoodensis Cockerell, 1905 ND-SD Rare .......XXX..Asteraceae: Chrysothamnus Nutt., Dieteria Nutt., Ericameria Nutt., Grindelia Willd., Helianthus L., Senecio L., Symphyotrichum Nees, Verbesina L.
Melissodes (Eumelissodes) grindeliae Cockerell, 1898 MN-ND-SD-TX Rare .....XXX....Asteraceae: Coreopsis L., Erigeron L., Grindelia Willd., Heterotheca Cass., Hymenoxys Cass., Ratibida Raf., Senecio L.
Melissodes (Eumelissodes) humilior Cockerell, 1903 TX Uncommon-Rare ......XX....Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Dieteria Nutt., Gutierrezia Lag., Helenium L., Helianthus L., Heterotheca Cass., Isocoma Nutt., Senecio L., Viguiera Kunth
Melissodes (Eumelissodes) illatus Lovell & Cockerell, 1906 MI-MN-MO-TX-WI Rare ......XXX... Asteraceae: Baccharis L., Bidens L., Centaurea L., Cirsium Mill., Coreopsis L., Cosmos Cav., Erigeron L., Gaillardia Foug., Grindelia Willd., Haplopappus Cass., Helenium L., Helianthus L., Heliopsis Pers., Hieracium L., Ratibida Raf., Rudbeckia L., Silphium L., Verbesina L., Vernonia Schreb., Xanthisma DC.
Melissodes (Eumelissodes) limbus LaBerge, 1961 TX Uncommon-Rare .......XX...Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Flaveria Juss., Gutierrezia Lag., Helianthus L., Heterotheca Cass., Hymenoxys Cass., Pectis L., Pityopsis Nutt., Solidago L., Symphyotrichum Nees, Verbesina L., Xanthisma DC.
Melissodes (Callimelissodes) lupinus Cresson, 1878 TX Rare ......XXXX..Asteraceae: Bidens L., Centaurea L., Cirsium Mill., Encelia Adans., Ericameria Nutt., Erigeron L., Grindelia Willd., Helianthus L., Hemizonia DC., Heterotheca Cass., Holocarpha Greene, Solidago L., Stephanomeria Nutt., Symphyotrichum Nees
Melissodes (Callimelissodes) lustrus LaBerge, 1961 IL Rare ......XX....Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Grindelia Willd., Isocoma Nutt., Symphyotrichum Nees
Melissodes (Eumelissodes) lutulentus LaBerge, 1961 KS-ND Rare .....XXXX... Asteraceae: Chrysothamnus Nutt., Grindelia Willd., Gutierrezia Lag., Heterotheca Cass.
Melissodes (Eumelissodes) manipularis Smith, 1854 IL Rare .......XX... Asteraceae: Chrysopsis (Nutt.) Elliot
Melissodes (Eumelissodes) menuachus Cresson, 1868 IA-IL-KS-MN-ND-NE-SD-TX-WI Uncommon-Rare ......XXXX..Asteraceae: Dieteria Nutt., Grindelia Willd., Erigeron L., Helianthus L., Heterotheca Cass., Isocoma Nutt., Machaeranthera Nees, Solidago L., Symphyotrichum Nees, Verbesina L., Viguiera Kunth
Melissodes (Eumelissodes) microstictus Cockerell, 1905 IL-TX Uncommon ......XXX...Asteraceae: Achillea L., Anaphalis DC., Antennaria Gaertn., Centaurea L., Chrysothamnus Nutt., Cirsium Mill., Ericameria Nutt., Erigeron L., Grindelia Willd., Helenium L., Helianthus L., Heterotheca Cass., Rudbeckia L., Senecio L., Solidago L., Symphyotrichum Nees
Melissodes (Eumelissodes) montanus Cresson, 1878 TX Rare .....XXXX...Asteraceae: Bidens L., Chrysothamnus Nutt., Coreopsis L., Erigeron L., Gymnosperma Less., Haplopappus Cass., Heliomeris Nutt., Heliopsis Pers., Machaeranthera Nees, Porophyllum Adans., Simsia Pers., Verbesina L., Viguiera Kunth, Xanthocephalum Willd, Xylothamia G.L. Nesom, Suh, D. Morgan & Simpson
Melissodes (Callimelissodes) nigracauda LaBerge, 1961 TX Rare .....XXXX... Asteraceae: Stephanomeria Nutt.
Melissodes (Eumelissodes) niveus Robertson, 1895 AR-IL-IN-KS-MO-NE-OH Uncommon ......XXXX.. Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Helianthus L., Solidago L., Symphyotrichum Nees, Vernonia Schreb.
Melissodes (Eumelissodes) pallidisignatus Cockerell, 1905 KS-ND-NE Rare ......XXXX..Asteraceae: Centaurea L., Coreopsis L., Engelmannia A. Gray ex Nutt., Ericameria Nutt., Erigeron L., Grindelia Willd., Gutierrezia Lag., Machaeranthera Nees, Senecio L., Solidago L., Symphyotrichum Nees
Melissodes (Eumelissodes) perlusus Cockerell, 1925 ND-NE Rare ......XX.... Asteraceae: Grindelia Willd., Helianthus L., Ratibida Raf.
Melissodes (Callimelissodes) plumosus LaBerge, 1961 ND Rare ....XXX.....Asteraceae: Centaurea L., Dieteria Nutt., Grindelia Willd., Helianthus L.
Melissodes (Heliomelissodes) rivalis Cresson, 1872 KS-MN-ND-NE-SD-TX Common .....XXX....Asteraceae: Brickellia Elliot, Centaurea L., Cirsium Mill., Helianthus L.
Melissodes (Eumelissodes) saponellus Cockerell, 1908 TX Rare ...XXXXXX...Asteraceae: Dieteria Nutt., Erigeron L., Symphyotrichum Nees
Melissodes (Eumelissodes) semilupinus Cockerell, 1905 MI-ND-TX Rare ......XXX...Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Gutierrezia Lag., Helianthus L., Machaeranthera Nees, Senecio L., Symphyotrichum Nees
Melissodes (Eumelissodes) snowii Cresson, 1878 ND-NE Rare ......XXX...Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Dieteria Nutt., Ericameria Nutt., Gaillardia Foug., Grindelia Willd., Gutierrezia Lag., Helianthus L., Heterotheca Cass., Isocoma Nutt., Senecio L., Stephanomeria Nutt., Symphyotrichum Nees, Xanthisma DC.
Melissodes (Eumelissodes) subagilis Cockerell, 1905 IA-KS-MN-ND-NE-SD-TX Uncommon-Rare .......XXX..Asteraceae: Chrysothamnus Nutt., Dieteria Nutt., Grindelia Willd., Helianthus L., Heterotheca Cass., Isocoma Nutt., Machaeranthera Nees, Pectis L., Symphyotrichum Nees
Melissodes (Eumelissodes) subillatus LaBerge, 1961 IL-IN-MI-MN-MO-ND-OH Uncommon .....XXXX... Asteraceae: Coreopsis L., Cirsium Mill., Rudbeckia L., Symphyotrichum Nees, Vernonia Schreb.
Melissodes (Eumelissodes) submenuacha Cockerell, 1897 TX Rare .......XXX.. Asteraceae: Chrysothamnus Nutt., Encelia Adans., Helianthus L., Heliopsis Pers., Verbesina L.
Melissodes (Eumelissodes) tinctus LaBerge, 1961 AR-IL-IN-MI-MN-MO-OH-TN-TX-WI Rare .......XXXXX Asteraceae: Chrysopsis (Nutt.) Elliot, Helianthus L., Symphyotrichum Nees, Verbesina L.
Melissodes (Eumelissodes) trinodis Robertson, 1901 AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-SD-TN-TX-WI Uncommon ......XXXX..Asteraceae: Coreopsis L., Grindelia Willd., Helianthus L., Heliopsis Pers., Ratibida Raf., Rudbeckia L., Solidago L., Symphyotrichum Nees
Melissodes (Eumelissodes) utahensis LaBerge, 1961 KS-NE-SD Uncommon-Rare ......XXX...Asteraceae: Chrysothamnus Nutt., Dieteria Nutt., Ericameria Nutt., Grindelia Willd., Gutierrezia Lag., Helianthus L., Solidago L., Symphyotrichum Nees, Verbesina L.
Melissodes (Eumelissodes) verbesinarum Cockerell, 1905 SD-TX Rare ....XXXXXXX.Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chaetopappa DC., Chrysothamnus Nutt., Dyssodia Cav., Encelia Adans., Grindelia Willd., Gutierrezia Lag., Helenium L., Helianthus L., Heterotheca Cass., Hymenoxys Cass., Isocoma Nutt., Machaeranthera Nees, Pectis L., Symphyotrichum Nees, Verbesina L., Xanthisma DC.
Melissodes (Eumelissodes) vernoniae Robertson, 1902 AR-IA-IL-IN-KS-MO-ND-NE-SD-TX Rare ....XXXXX... Asteraceae: Bidens L., Gaillardia Foug., Grindelia Willd., Helianthus L., Heliopsis Pers., Lactuca L., Liatris Gaertn. ex Schreb., Rudbeckia L., Silphium L., Vernonia Schreb.
Melissodes (Eumelissodes) wheeleri Cockerell, 1906 AR-IL-IN-KS-MI-MN-MO-ND-NE-OK-TN-TX-WI Rare ...XXXXXXX..Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Coreopsis L., Echinacea Moench, Engelmannia A. Gray ex Nutt., Gaillardia Foug., Grindelia Willd., Helianthus L., Pectis L., Ratibida Raf., Rudbeckia L., Silphium L., Verbesina L.
Apidae: Apinae: Eucerini Peponapis (Peponapis) pruinosa (Say, 1837) IA-IL-IN-KS-MI-MN-MO-ND-NE-OK-SD-TN-TX-WI Common .....XXXX... Cucurbita L.
Peponapis (Xeropeponapis) timberlakei Hurd & Linsley, 1964 MO Rare ......XXX... Cucurbita L.
Peponapis (Eopeponapis) utahensis (Cockerell, 1905) TX Rare .......XXXXX Cucurbita L.
Apidae: Apinae: Eucerini Svastra (Epimelissodes) aegis (LaBerge, 1956) AR-TX Rare ......XXXX.. Asteraceae: Balduina Nutt., Chrysopsis (Nutt.) Elliot, Helenium L., Helianthus L., Symphyotrichum Nees
Svastra (Anthedonia) compta (Cresson, 1878) IL-IN-KS-MO-NE-OK-TN-TX Rare .....XXXXXX. Oenothera L.
Svastra (Idiomelissodes) duplocincta (Cockerell, 1905) TX Rare ......XXX... Ferocactus Britton & Rose
Svastra (Epimelissodes) helianthelli (Cockerell, 1905) KS-TX Rare ......XX.... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Helianthus L.
Svastra (Epimelissodes) machaerantherae (Cockerell, 1904) TX Rare ......XXXX.. Asteraceae: Baccharis L., Ericameria Nutt., Helianthus L., Heterotheca Cass., Verbesina L.
Svastra (Epimelissodes) obliqua (Say, 1837) AR-IA-IL-IN-KS-KY-MI-MN-MO-ND-NE-OH-OK-SD-TN-TX-WI Common-Uncommon .....XXXXXX. Asteraceae: Borrichia Adans., Chrysothamnus Nutt., Cirsium Mill., Grindelia Willd., Helianthus L., Rudbeckia L., Vernonia Schreb.
Svastra (Epimelissodes) petulca (Cresson, 1878) AR-IL-IN-KS-KY-MO-NE-OK-TN-TX Common-Uncommon ...XXXXXX... Asteraceae: Coreopsis L., Helenium L., Helianthus L., Pyrrhopappus DC., Ratibida Raf., Rudbeckia L., Verbesina L.
Svastra (Epimelissodes) sabinensis (Cockerell, 1924) TX Uncommon ......XXXX.. Asteraceae: Baccharis L., Baileya Harv. & A. Gray ex A. Gray, Bebbia (Benth.) Greene, Encelia Adans., Gaillardia Foug., Haplopappus Cass., Helianthus L., Heterotheca Cass., Hymenothrix A. Gray, Isocoma Nutt.
Svastra (Epimelissodes) sila (LaBerge, 1956) TX Rare ......XXX... Psilostrophe DC.
Svastra (Epimelissodes) texana (Cresson, 1872) AR-KS-OK-TX Uncommon-Rare ......XXXX.. Asteraceae: Chrysothamnus Nutt., Verbesina L.
Apidae: Apinae: Eucerini Syntrichalonia exquisita (Cresson, 1878) TX Rare .......XXXX. Asteraceae: Helianthus L., Rudbeckia L., Verbesina L., Viguiera Kunth
Apidae: Apinae: Eucerini Tetraloniella (Pectinapis) fasciatella (LaBerge, 1970) TX Rare .....X...... Salvia L.
Tetraloniella (Tetraloniella) lippiae (Cockerell, 1904) TX Rare .......XX... Sphaeralcea A. St.-Hil.
Apidae: Apinae: Eucerini Xenoglossa (Eoxenoglossa) kansensis Cockerell, 1905 IA-KS-MN-NE-OK-TX-WI Rare ....XXXX.... Cucurbita L., Ipomoea L.
Xenoglossa (Xenoglossa) patricia Cockerell, 1896 TX Rare ......XX.... Cucurbita L.
Xenoglossa (Eoxenoglossa) strenua (Cresson, 1878) IA-IL-IN-KS-NE-OH-OK-TX-WI Uncommon .....XXX.... Cucurbita L.
Apidae: Apinae: Exomalopsini Anthophorula (Anthophorisca) asteris (Mitchell, 1962) AR-IN-TX Rare .......XX... Symphyotrichum Nees
Anthophorula (Anthophorisca) micheneri Timberlake, 1948 KY-TX Rare ........XX.. Agalinis Raf.
Apidae: Apinae: Exomalopsini Exomalopsis (Stilbomalopsis) solidaginis Cockerell, 1989 TX Rare .....XXXXX.. Asteraceae: Baccharis L., Baileya Harv. & A. Gray ex A. Gray, Gutierrezia Lag., Helianthus L., Heterotheca Cass., Machaeranthera Nees, Pectis L.
Colletidae Caupolicana ocellata Michener, 1966 KS-TX Rare .....XXXX... Dalea L., Psorothamnus Rydb.
Colletidae Colletes aberrans Cockerell, 1897 IA-IL-IN-KS-MI-ND-NE-SD-TX-WI Rare ......XX.... Dalea L.
Colletes aestivalis Patton, 1879 IL-IN-MI-MO-OH-TN Rare ....XXX..... Heuchera L. [Boykinia Nutt., Mitella L., Tiarella L.?]
Colletes albescens Cresson, 1868 IL-KS-ND-NE-SD-WI Rare ...XXXX..... Amorpha L., Dalea L.
Colletes algarobiae Cockerell, 1900 TX Rare .XXXX....... Prosopis L.
Colletes americanus Cresson, 1868 AR-IA-IL-IN-KS-MI-MO-ND-NE-OH-SD-TX-WI Uncommon ....XXXXXX.. Asteraceae: Euthamia Nutt. ex Cass., Grindelia Willd., Helianthus L., Pityopsis Nutt., Rudbeckia L., Solidago L., Symphyotrichum Nees
Colletes andrewsi Cockerell, 1906 ND-NE-SD-WI Rare ....XXX..... Heuchera L.
Colletes banksi Swenk, 1908 MI-TN Rare ...XXXX..... Ilex L.
Colletes birkmanni Swenk, 1906 KS-OK-TX Rare ...XXX..... Asteraceae: Baccharis L., Gutierrezia Lag., Helianthus L., Heliopsis Pers., Hymenopappus L'Hér.
Colletes brevicornis Robertson, 1897 IL-IN-KS-MI-MN-MO-ND-NE-OK-SD-TX-WI Uncommon-Rare ..XXXX...... Campanula L., Triodanis Raf. ex Greene
Colletes bumeliae Neff, 2004 TX Rare ...XXX...... Sideroxylon L.
Colletes ciliatus Patton, 1879 IA-IL-KS-NE Rare ........XX.. Cuscuta L.
Colletes clypeonitens Swenk, 1906 TX Uncommon-Rare ..XX........ Larrea Cav.
Colletes compactus Cresson, 1868 IA-IL-IN-KS-KY-MI-MN-OH-TN-TX-WI Uncommon-Rare .....XXXXXX. Asteraceae: Bidens L., Chrysopsis (Nutt.) Elliot, Chrysothamnus Nutt., Ericameria Nutt., Rudbeckia L., Solidago L., Symphyotrichum Nees
Colletes fulgidus Swenk, 1904 SD-TX Rare ......XXX... Asteraceae: Achillea L., Chrysothamnus Nutt., Cirsium Mill., Ericameria Nutt., Erigeron L., Grindelia Willd., Gutierrezia Lag., Haplopappus Cass., Rudbeckia L., Solidago L.
Colletes gypsicolens Cockerell, 1897 TX Rare ........XX.. Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Grindelia Willd., Gutierrezia Lag., Helianthus L., Verbesina L.
Colletes inuncantipedis Neff, 2004 TX Rare .....X...... Sideroxylon L.
Colletes laticinctus Timberlake, 1951 NE Rare ......XXXX.. Asteraceae: Gutierrezia Lag.
Colletes latitarsis Robertson, 1891 AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-OK-SD-TN-TX-WI Uncommon ..XXXXXXX... Physalis L.
Colletes petalostemonis Swenk, 1906 ND-NE-SD Rare ......X..... Fabaceae: Dalea L., Psorothamnus Rydb.
Colletes productus Robertson, 1891 IL-MO-TN Rare ...XXXX..... Lyonia Nutt., Vaccinium L.: Vaccinium arboreum Marshall
Colletes prosopidis Cockerell, 1897 TX Uncommon-Rare ..XXX....... Prosopis L.
Colletes robertsonii Dalla Torre, 1896 IA-IL-KS-MN-MO-ND-NE-OK-SD Rare .....XXX.... Dalea L.
Colletes rufocinctus Cockerell, 1929 MN-ND-SD Rare .......X.... Asteraceae: Grindelia Willd., Gutierrezia Lag., Helianthus L., Ratibida Raf., Solidago L., Symphyotrichum Nees
Colletes scopiventer Swenk, 1908 OK-TX Rare .......XX... Chamaesaracha (A. Gray) Benth.
Colletes simulans Cresson, 1868 AR-IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-SD-TN-TX-WI Common-Uncommon ....XXXXXXX. Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Euthamia Nutt. ex Cass., Solidago L., Symphyotrichum Nees
Colletes solidaginis Swenk, 1906 IA-IL-IN-MI-MN-ND-NE Rare ......XXX... Solidago L.
Colletes speculiferus Cockerell, 1927 IL-MN-NE-TX Rare ......XXXX.. Asteraceae: Euthamia Nutt. ex Cass., Solidago L., Symphyotrichum Nees
Colletes sphaeralceae Timberlake, 1951 TX Rare ...XXX...... Sphaeralcea A. St.-Hil.
Colletes susannae Swenk, 1925 IA-KS-MN-ND-NE-SD-WI Rare ......XX.... Amorpha L., Dalea L.
Colletes validus Cresson, 1868 IL-IN-MI-WI Rare ..XXXXX..... Ericaceae: Vaccinium L.
Colletes willistoni Robertson, 1891 IL-KS-MI-MO-ND-NE-OH-SD-TX-WI Rare ...XXXXXX... Physalis L.
Colletes wilmattae Cockerell, 1904 IA-MN-ND-NE-TX-WI Rare .....XX..... Dalea L.
Colletes xerophilus Timberlake, 1951 TX Rare ...XXX...... Salix L.
Halictidae: Halictinae: Halictini Lasioglossum (Sphecodogastra) aberrans (Crawford, 1903) ND-NE-SD-TX Rare ....XXXXXX.. Oenothera L.
Lasioglossum (Sphecodogastra) lusorium (Cresson, 1872) KS-ND-NE-SD-TX-WI Rare ..XXXXXXXX.. Oenothera L.
Lasioglossum (Hemihalictus) lustrans (Cockerell, 1897) AR-IN-KS-KY-MI-ND-OH-OK-SD-TN-TX-WI Rare ...XXXXXX... Pyrrhopappus DC.
Lasioglossum (Sphecodogastra) noctivaga Linsley & MacSwain, 1962 KS-OK-TX Rare ..XXXXXXXX.. Oenothera L.
Lasioglossum (Sphecodogastra) oenotherae (Stevens, 1920) IL-KS-MI-TX Rare .....XXX.... Oenothera L.
Lasioglossum (Hemihalictus) pectinatum (Robertson, 1890) IL-KY-OH Rare .....XXX.... Physalis L.
Lasioglossum (Sphecodogastra) texanum (Cresson, 1872) IA-IL-IN-KS-MI-MN-MO-ND-NE-OK-SD-TX-WI Rare ...XXXXXXXXX Oenothera L.
Halictidae: Nomiinae Dieunomia (Dieunomia) heteropoda (Say, 1824) AR-IA-IL-IN-KS-MI-MN-MO-NE-OH-OK-TN-TX-WI Uncommon-Rare ......XXX... Asteraceae: Bidens L., Coreopsis L., Helianthus L., Silphium L., Solidago L., Verbesina L.
Dieunomia (Epinomia) nevadensis (Cresson, 1874) AR-IA-IL-KS-OK-TN-TX Uncommon-Rare .......XX... Asteraceae: Baccharis L., Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Gaillardia Foug., Grindelia Willd., Gutierrezia Lag., Helianthus L., Heterotheca Cass., Pectis L., Verbesina L.
Halictidae: Rophitinae Conanthalictus cotullensis Crawford, 1907 TX Rare ...XX....... Nama L.
Conanthalictus bakeri Crawford, 1907 TX Rare ..XXXX...... Phacelia Juss.
Halictidae: Rophitinae Dufourea marginata (Cresson, 1878) IA-IL-KS-MN-ND-NE-SD-WI Uncommon .......XX... Asteraceae: Chrysothamnus Nutt., Grindelia Willd., Helianthus L., Solidago L.
Dufourea monardae (Viereck, 1924) IA-IL-IN-MI-MN-WI-TN Rare ......XX.... Monarda L.
Dufourea novaeangliae (Robertson, 1897) IL-MI-MO-OH-WI Uncommon-Rare ....XXX..... Pontederia L.
Dufourea pulchricornis (Cockerell, 1916) TX Uncommon-Rare ..XX........ Brassicaceae: Lepidium L.
Halictidae: Rophitinae Sphecodosoma (Michenerula) beameri (Bohart, 1965) TX Rare .XXX........ Nama L.
Sphecodosoma (Sphecodosoma) dicksoni (Timberlake, 1961) TX Rare ..XXXXXX.... Nama L.
Megachilidae: Lithurginae Lithurgopsis apicalis (Cresson, 1875) KS-ND-NE-OK-SD-TX Uncommon-Rare ...XXX...... Cactaceae: Cylindropuntia (Engelm.) Kreuzinger, Echinocactus Link & Otto, Ferocactus Britton & Rose, Opuntia Mill.
Lithurgopsis gibbosa (Smith, 1853) KS-OK-TX Rare ...XX....... Opuntia Mill.
Lithurgopsis littoralis (Cockerell, 1917) IL-KS-OK-TX Rare ...XX....... Cactaceae: Cylindropuntia (Engelm.) Kreuzinger, Echinocactus Link & Otto, Ferocactus Britton & Rose, Opuntia Mill.
Megachilidae: Megachilinae: Anthidiini Anthidium (Anthidium) atripes Cresson, 1879 TX Rare ...XXX...... Fabaceae: Astragalus L., Lotus L.
Anthidium (Anthidium) clypeodentatum Swenk, 1914 ND-NE-SD Rare .....XXX.... Fabaceae: Astragalus L., Lotus L.
Anthidium (Anthidium) emarginatum (Say, 1824) KS-NE-TX Uncommon-Rare ...XXXXX.... Phacelia Juss.
Anthidium (Anthidium) mormonum Cresson, 1878 NE Uncommon-Rare ...XXXXXX... Lotus L., Phacelia Juss.
Anthidium (Anthidium) palmarum Cockerell, 1904 TX Uncommon-Rare ..XXXX...... Phacelia Juss.
Anthidium (Anthidium) tenuiflorae Cockerell, 1907 MI-MN-ND-NE-SD Uncommon-Rare ....XXXX.... Fabaceae: Astragalus L., Lotus L.
Anthidium (Anthidium) utahense Swenk, 1914 NE Rare ....XXXXXX.. Fabaceae: Lotus L.
Megachilidae: Megachilinae: Anthidiini Dianthidium (Dianthidium) curvatum (Smith, 1854) AR-KS-MO-ND-NE-SD-TX Uncommon-Rare ...XXXXXX... Asteraceae: Coreopsis L., Grindelia Willd., Helenium L., Helianthus L.
Dianthidium (Dianthidium) parvum (Cresson, 1878) TX Rare .....XXXXX.. Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Cirsium Mill., Encelia Adans., Erigeron L., Gaillardia Foug., Grindelia Willd., Gutierrezia Lag., Heterotheca Cass., Machaeranthera Nees, Stephanomeria Nutt.
Dianthidium (Dianthidium) platyurum Cockerell, 1923 TX Rare .....XXXXX.. Asteraceae: Bebbia (Benth.) Greene, Erigeron L., Gutierrezia Lag., Heterotheca Cass., Malacothrix DC., Pectis L., Senecio L., Stephanomeria Nutt.
Dianthidium (Dianthidium) simile (Cresson, 1864) IN-MI-MN-WI Rare ......XXX... Asteraceae: Solidago L., Symphyotrichum Nees
Dianthidium (Dianthidium) ulkei (Cresson, 1878) ND-NE-SD Uncommon-Rare .....XXXXX.. Asteraceae: Achillea L., Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Cirsium Mill., Dieteria Nutt., Encelia Adans., Ericameria Nutt., Erigeron L., Grindelia Willd., Gutierrezia Lag., Haplopappus Cass., Helenium L., Helianthus L., Heterotheca Cass., Machaeranthera Nees, Solidago L., Stephanomeria Nutt., Symphyotrichum Nees, Verbesina L.,
Megachilidae: Megachilinae: Anthidiini Paranthidium (Paranthidium) jugatorium (Say, 1824) AR-IA-IL-KY-MN-NE-OH-TN Uncommon-Rare ......XXX... Asteraceae: Erigeron L., Grindelia Willd., Haplopappus Cass., Helianthus L., Heterotheca Cass., Rudbeckia L., Silphium L., Verbesina L., Viguiera Kunth
Megachilidae: Megachilinae: Anthidiini Trachusa (Heteranthidium) bequaerti (Schwarz, 1926) TX Rare .....XXX.... Dalea L.
Trachusa (Heteranthidium) larreae (Cockerell, 1897) TX Uncommon-Rare ..XXX....... Larrea Cav.
Trachusa (Heteranthidium) zebrata (Cresson, 1872) IN-MI-MN-NE-SD-TX-WI Rare .......XXX.. Asteraceae: Encelia Adans., Helianthus L.
Megachilidae: Megachilinae: Megachilini Mechachile (Phaenosarus) fortis Cresson, 1872 IA-IL-IN-KS-MN-ND-NE-OK-SD-TX-WI Rare ......XXX... Helianthus L.
Megachile (Phaenosarus) agustini Cockerell, 1905 TX Rare ......XXX... Asteraceae: Cirsium Mill., Helianthus L., Heliomeris Nutt., Verbesina L.
Megachile (Chelostomoides) chilopsidis Cockerell, 1900 TX Uncommon-Rare ..XXX....... Fabaceae: Parkinsonia L., Prosopis L., Psorothamnus Rydb.
Megachile (Sayapis) fidelis Cresson, 1878 KS-NE-SD Rare .....XXX.... Asteraceae: Centaurea L., Chrysothamnus Nutt., Cirsium Mill., Coreopsis L., Dieteria Nutt., Encelia Adans., Ericameria Nutt., Erigeron L., Grindelia Willd., Gutierrezia Lag., Haplopappus Cass., Helenium L., Helianthus L., Heterotheca Cass., Madia Molina, Rudbeckia L., Senecio L., Solidago L., Symphyotrichum Nees, Verbesina L., Viguiera Kunth
Megachile (Sayapis) inimica Cresson, 1872 AR-IA-IL-IN-KS-KY-MI-MO-NE-OH-TN-TX Uncommon ......XXXXX. Asteraceae: Baccharis L., Chrysopsis (Nutt.) Elliot, Cirsium Mill., Coreopsis L., Erigeron L., Grindelia Willd., Helianthus L., Rudbeckia L., Silphium L., Solidago L., Verbesina L., Vernonia Schreb.
Megachile (Megachiloides) integra Cresson, 1878 IL-IN-KS-OH-SD-TX Rare ..XXXXXXX... Galactia P. Br., Strophostyles Elliot
Megachile (Chelostomoides) lobatifrons Cockerell, 1924 TX Rare ....XXXXXX.. Fabaceae: Parkinsonia L., Prosopis L., Psorothamnus Rydb.
Megachile (Megachiloides) manifesta Cresson, 1878 ND-NE Rare .....XXXX... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Erigeron L., Grindelia Willd., Helianthus L., Heliomeris Nutt., Senecio L., Xanthisma DC.
Megachile (Xanthosarus) melanophaea Smith, 1853 MI-MN-ND-NE-SD-TX-WI Uncommon ....XXXX.... Fabaceae: Astragalus L., Hedysarum L., Lotus L., Lupinus L.
Megachile (Sayapis) newberryae Cockerell, 1900 TX Uncommon-Rare ..XXX....... Fabaceae: Dalea L., Parkinsonia L., Prosopis L.
Megachile (Chelostomoides) odontostoma Cockerell, 1924 TX Rare ..XXX....... Fabaceae: Parkinsonia L., Prosopis L.
Megachile (Megachiloides) oenotherae (Mitchell, 1924) OK-TX Rare ...XXX...... Oenothera L.
Megachile (Argyropile) parallela Smith, 1853 AR-IA-IL-IN-KS-MO-ND-NE-OH-OK-SD-TN-TX-WI Common-Uncommon .....XXXX... Asteraceae: Baccharis L., Bebbia (Benth.) Greene, Centaurea L., Chrysothamnus Nutt., Coreopsis L., Echinacea Moench, Ericameria Nutt., Gaillardia Foug., Grindelia Willd., Gutierrezia Lag., Haplopappus Cass., Helenium L., Helianthus L., Heterotheca Cass., Hymenothrix A. Gray, Malacothrix DC., Palafoxia Lag., Pyrrocoma Hook., Rudbeckia L., Senecio L., Solidago L., Stephanomeria Nutt., Tetraneuris Greene, Verbesina L.
Megachile (Xanthosarus) perihirta Cockerell, 1898 IA-IL-KS-ND-NE-SD-TX Uncommon-Rare ....XXXXX... Asteraceae?, Fabaceae?
Megachile (Sayapis) pugnata Say, 1837 IA-IL-IN-KS-MI-MN-MO-ND-NE-OH-SD-TN-TX-WI Uncommon .....XXXX... Asteraceae: Cirsium Mill., Coreopsis L., Erigeron L., Grindelia Willd., Helianthus L., Rudbeckia L..
Megachile (Argyopile) rossi Mitchell, 1943 TX Uncommon-Rare ......XX.... Asteraceae: Encelia Adans., Helianthus L., Pluchea Cass., Verbesina L.
Megachile (Megachiloides) soledadensis Cockerell, 1900 TX Rare ......XXX... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Bebbia (Benth.) Greene, Chrysothamnus Nutt., Heliomeris Nutt., Helianthus L., Heterotheca Cass., Machaeranthera Nees, Pectis L., Verbesina L.
Megachile (Chelostomoides) subexilis Cockerell, 1908 NE-SD-TX Uncommon-Rare ......XXX... Fabaceae: Lotus L., Lupinus L.
Megachile (Megachiloides) subnigra Cresson, 1879 TX Rare ...XXXX..... Asteraceae: Achillea L., Bebbia (Benth.) Greene, Chaenactis DC., Cirsium Mill., Coreopsis L., Encelia Adans., Ericameria Nutt., Eriophyllum Lag., Senecio L.
Megachile (Argyropile) townsendiana Cockerell, 1898 OK-TX Uncommon-Rare .....XXXX... Asteraceae: Baccharis L., Baileya Harv. & A. Gray ex A. Gray, Chrysothamnus Nutt., Encelia Adans., Haplopappus Cass., Heterotheca Cass., Hymenopappus L'Hér., Isocoma Nutt., Pectis L., Verbesina L.
Megachile (Megachiloides) wheeleri Mitchell, 1927 ND-NE Rare ......XXX... Asteraceae: Achillea L., Chrysothamnus Nutt., Cirsium Mill., Ericameria Nutt., Erigeron L., Grindelia Willd., Machaeranthera Nees,
Megachile (Melanosarus) xylocopoides Smith, 1853 AR-IL-KY-MO-OH-OK-TN-TX Uncommon ..XXXXXXXX.. Asteraceae: Coreopsis L., Erigeron L., Helianthus L., Rudbeckia L., Solidago L., Vernonia Schreb.
Megachilidae: Megachilinae: Osmiini Ashmeadiella (Ashmeadiella) bucconis (Say, 1837) AR-IL-IN-KS-MI-MO-ND-NE-SD-TX-WI Uncommon-Rare ....XXXX.... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chaenactis DC., Chrysothamnus Nutt., Coreopsis L., Dieteria Nutt., Echinacea Moench, Encelia Adans., Ericameria Nutt., Erigeron L., Grindelia Willd., Gutierrezia Lag., Heliomeris Nutt., Heterotheca Cass., Hymenothrix A. Gray, Isocoma Nutt., Lactuca L., Pectis L., Senecio L., Solidago L., Verbesina L., Xanthisma DC.
Ashmeadiella (Ashmeadiella) cubiceps (Cresson, 1979) TX Rare ...XXX...... Asteraceae: Baileya Harv. & A. Gray ex A. Gray, Chaenactis DC., Encelia Adans., Ericameria Nutt., Erigeron L., Heliomeris Nutt., Hieracium L., Malacothrix DC.
Ashmeadiella (Arogochila) erema Michener, 1939 TX Rare ...XXX...... Fabaceae: Dalea L., Hedysarum L., Lupinus L., Psorothamnus Rydb.
Ashmeadiella (Ashmeadiela) opuntiae (Cockerell, 1897) SD-TX Uncommon-Rare ...XXXXXX... Cactaceae: Cylindropuntia (Engelm.) Kreuzinger, Echinocactus Link & Otto, Ferocactus Britton & Rose, Opuntia Mill.
Ashmeadiella (Ashmeadiella) prosopidis (Cockerell, 1897) TX Uncommon-Rare ..XX........ Fabaceae: Dalea L., Parkinsonia L., Prosopis L., Psorothamnus Rydb.
Ashmeadiella (Chilosima) rhodognatha Cockerell, 1924 TX Rare ..XXX....... Fabaceae: Dalea L., Parkinsonia L., Prosopis L., Psorothamnus Rydb.
Ashmeadiella (Isosmia) rubrella (Michener, 1949) TX Rare ..XXX....... Fabaceae: Dalea L., Prosopis L., Psorothamnus Rydb.
Megachilidae: Megachilinae: Osmiini Atoposmia (Eremosmia) daleae (Michener, 1951) TX Rare ..XXXXXX.... Dalea L.
Megachilidae: Megachilinae: Osmiini Heriades (Neotrypetes) cressoni Michener, 1938 NE Rare ......XX.... Asteraceae: Chrysothamnus Nutt., Ericameria Nutt., Erigeron L., Grindelia Willd., Helianthus L., Machaeranthera Nees, Senecio L.
Megachilidae: Megachilinae: Osmiini Hoplitis (Alcidamea) biscutellae (Cockerell, 1897) TX Rare ..XXXX...... Larrea Cav.
Hoplitis (Proteriades) incanescens (Cockerell, 1922) SD Rare .....X...... Cryptantha Lehm. ex G. Don
Hoplitis (Robertsonella) micheneri Mitchell, 1962 KS-MO Rare ...XX....... Amorpha L.
Hoplitis (Robertsonella) simplex (Cresson, 1864) AR-IL-KS-MI-MO-OH-OK-TN-TX Rare ..XXX....... Nemophila Nutt., Phacelia Juss.
Megachilidae: Megachilinae: Osmiini Osmia (Cephalosmia) californica Cresson, 1864 SD Rare ....XXX..... Asteraceae: Balsamorhiza Nutt., Chaenactis DC., Cirsium Mill., Ericameria Nutt., Heliomeris Nutt., Heterotheca Cass., Senecio L.
Osmia (Helicosmia) chalybea Smith, 1853 AR-OH-TX Rare ..XXX....... Cirsium Mill.
Osmia (Helicosmia) coloradensis Cresson, 1878 AR-KS-NE-TN-TX Uncommon-Rare ...XXXXX.... Asteraceae: Chaenactis DC., Cirsium Mill., Coreopsis L., Encelia Adans., Erigeron L., Senecio L.
Osmia (Melanosmia) dakotensis Michener, 1937 ND-SD Rare ....XXXX.... Fabaceae: Astragalus L., Hedysarum L.
Osmia (Melanosmia) distincta Cresson, 1864 IL-IN-KS-MI-MO-ND-NE-OH-SD-TN Uncommon ..XXXXX..... Penstemon Schmidel
Osmia (Melanosmia) integra Cresson, 1978 MI-NE-SD-TX Rare ..XXXXX..... Fabaceae: Astragalus L., Lotus L., Lupinus L., Thermopsis R. Br.
Osmia (Trichinosmia) latisulcata Michener, 1936 TX Rare ...XX....... Fabaceae: Astragalus L., Lotus L., Lupinus L.
Osmia (Melanosmia) longula Cresson, 1864 SD Rare ...XXXXX.... Fabaceae: Astragalus L., Hedysarum L., Lupinus L.
Osmia (Cephalosmia) montana Cresson, 1864 KS-TX Rare ..XXXXXX.... Asteraceae: Agoseris Raf., Chaenactis DC., Cirsium Mill., Encelia Adans., Ericameria Nutt., Erigeron L., Eriophyllum Lag., Grindelia Willd., Helianthella Torr. & A. Gray, Senecio L.
Osmia (Melanosmia) nigrifrons Cresson, 1878 ND-SD Rare ...XXXXX.... Fabaceae: Astragalus L., Lupinus L., Trifolium L.
Osmia (Cephalosmia) subaustralis Cockerell, 1900 MI-SD Rare ...XXXXX.... Asteraceae: Chaenactis DC., Cirsium Mill., Encelia Adans., Erigeron L., Grindelia Willd., Helianthus L., Heliomeris Nutt., Senecio L.
Osmia (Helicosmia) texana Cresson, 1872 MI-OH-ND-NE-SD-TX Uncommon-Rare ..XXXXXX.... Cirsium Mill.
Osmia (Melanosmia) titusi Cockerell, 1905 TX Rare ..XX........ Fabaceae: Astragalus L., Dalea L., Lotus L., Lupinus L.
Osmia (Melanosmia) virga Sandhouse, 1939 IN-MI-MN-TN-WI Rare ...XXX...... Vaccinium L.
Melittidae: Dasypodainae: Hesperapini Hesperapis (Carinapis) carinata Stevens, 1919 IL-IN-KS-ND-NE Rare .......XX... Asteraceae: Helianthus L.
Hesperapis (Amblyapis) larreae Cockerell, 1907 TX Rare ..XXXX...... Larrea Cav.
Hesperapis (Carinapis) rhodocerata (Cockerell, 1897) TX Uncommon .......XX... Asteraceae: Bahia Lag., Baileya Harv. & A. Gray ex A. Gray, Chloracantha G.L. Nesom, Dyssodia Cav., Gutierrezia Lag., Helianthus L., Heliomeris Nutt., Heterotheca Cass., Hymenothrix A. Gray, Isocoma Nutt., Machaeranthera Nees, Pectis L., Verbesina L., Xylothamia G.L. Nesom, Suh, D. Morgan & Simpson
Hesperapis (Hesperapis) trochanterata Snelling, 1987 TX Rare ....XXXX.... Nama L.
Melittidae: Melittinae: Macropidini Macropis (Macropis) ciliata Patton, 1880 IL-OH-WI Rare ....XXXX.... Lysimachia L.
Macropis (Macropis) nuda (Provancher, 1882) IL-MI-MN-ND-SD-WI Uncommon-Rare .....XXXX... Lysimachia L.
Macropis (Macropis) patellata Patton, 1880 IA-IL-MO-NE Rare .....XXX.... Lysimachia L.
Macropis (Macropis) steironematis Robertson, 1891 IA-IL-KS-MO-MN-NE-TN Rare .....XX..... Lysimachia L.
Melittidae: Melittinae: Melittini Melitta (Cilissa) eickworti Snelling & Stage, 1995 TN Rare ....XX...... Vaccinium (Polycodium) L.
Melitta (Cilissa) melittoides (Viereck, 1909) TN Rare .....XX..... Lyonia Nutt.

Recommendations

Pollen specialist bees of the Central United States were most often mesolectic on Asteraceae (198 spp.), Fabaceae (29 spp.), Brassicaceae (9 spp.), Malvaceae (7 spp.), and Cactaceae (5 spp.) and monolectic on Salix L. (19 spp.), Oenothera L. (11 spp.), Phacelia Juss. (7 spp.), Physalis L. (7 spp.), Vaccinium L. (7 spp.), Cucurbita L. (6 spp.), Larrea (6 spp.), Chamaesaracha (A. Gray) Benth. (5 spp.), Cornus (Swida) L. (4 spp.), and Lysimachia L. (4 spp.). Twenty-three unique relationships occured between individual monoleges and single host plants. Host plants were often either community dominant species or had early/late season blooms, crepuscular blooms, and odd floral morphologies.

Over 60% of the pollen specialist bees listed were rare. Pollinator habitat conservation and enhancement projects should prioritize practices that foster diverse communities of rare native pollen specialist bees and associated host plants (Fowler 2016a, 2016b). Site-specific practices including beetle banks, companion plants, cover crops, field borders, hedgerows, insectaries, meadows, pastures, prairies, and riparian buffers could be composed primarily of host plants for pollen specialist bees and secondarily of pollen and nectar plants for generalist pollinators. Local native plant horticulturists and nurseries should prioritize the propagation of ecoregional host plants for pollen specialist bees.

Conservation practices can protect native pollen specialist bees from population declines and extinctions, while identically supporting other managed and wild beneficial insects, providing wildlife habitat, reducing weeds and erosion, improving soil health and water quality, regulating pests, offering harvestable products, and bettering aesthetics (Garibaldi et al. 2014; Kleijn et al. 2015; Wratten et al. 2012). Therefore, habitat conservation and enhancement for native pollen specialist bees works synergistically to promote strong environmental, social, and economic sustainability in the Central United States.

Acknowledgements

Thank you for comments, Sam Droege (USGS BIML)... Thanks, BugGuide, BONAP, Discover Life, and USDA-NRCS PLANTS. Thank you for taxonomy and state attributions, John Ascher (National University of Singapore). Thanks for decriptions, authors: Cockerell, Cresson, Timberlake, Robertson, LaBerge, Viereck, Smith, Mitchell, Swenk, Say, Crawford, Viereck & Cockerell, Michener, Provancher, Swenk & Cockerell, Neff, Patton, Ribble, Cockerell & Porter, Stevens, Kirby, Dalla Torre, Linsley & MacSwain, Casad, Ashmead, Shinn, Shinn & Engel, Fabricius, Neff & Larkin, Gusenleitner & Schwartz, Donovan, Lanham, LaBerge & Ribble, Casad & Cockerell, Rozen, Portman & Griswold, Portman & Neff, Griswold, Lovell & Cockerell, Hurd & Linsley, Bohart, Schwarz, Sandhouse, Snelling, Snelling & Stage.

Thank you for sharing records, collectors: American Museum of Natural History, John Ascher, AMNH, Bee Biology and Systematics Laboratory, California State Collection of Arthropods, Computarización de la colección de abejas (Hymenoptera: Apoidea) del Museo de Zoología Alfonso L. Herrera, de la Facultad de Ciencias de la UNAM, CNIN/Abejas de México/Apoidea, Actualización de la base de datos del proyecto H278 Apoidea (Hymenoptera) del Valle de Zapotitlán de las Salinas, Puebla, Dartmouth University Bee Collection, Essig Museum Entomology Collection, Global Biodiversity Information Facility, Illinois Natural History Survey, iNaturalist, Los Angeles County Museum, Lund Museum of Zoology, Bob Minckley, NMNH Entomology Collections, Rocky Mountain Biological Laboratory Bee Collection, Rutgers University Arthropod Collection, Jessica Rykken, UAM Entomology Observations (Arctos), Snow Entomology Collection, University of Kansas, University of California Riverside Entomology Research Museum, United States Geological Survey, Sam Droege...

References

Records or requests? Email: j@jarrodfowler.com

Photos from The USGS Bee Inventory and Monitoring Lab

Visit: Pollen Specialist Bees of the Eastern United States

Visit: Pollen Specialist Bees of the Western United States

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